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Binomial nomenclature
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{{short description|System of identifying species of organisms using a two-part name}}{{redirect|Latin name|personal names in the Roman Empire|Roman naming conventions|the practice of rendering names in a Latin style|Latinisation of names}}{{use dmy dates|date=January 2018}}File:Killerwhales jumping.jpg|thumb|Orcinus orcaOrcinus orcaFile:Starr 070320-5799 Echinopsis pachanoi.jpg|thumb|Trichocereus macrogonus var. pachanoiTrichocereus macrogonus var. pachanoiIn taxonomy, binomial nomenclature (“two-term naming system“), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, although they can be based on words from other languages. Such a name is called a binomial name (which may be shortened to just “binomial“), a binomen, {{not a typo|binominal|reason=“Binomi’n’al”, with an “n” before the “al”, is the alternative name in the ICZN. Do not “correct” it.}} name, or a scientific name; more informally it is also historically called a Latin name. In the ICZN, the system is also called {{not a typo|binominal|reason=“Binomi’n’al”, with an “n” before the “al”, is the alternative name in the ICZN. Do not “correct” it.}} nomenclature, “binomi’N’al” with an “N” before the “al”, which is {{em|not}} a typographic error, meaning “two-name naming system”.The first part of the name – the generic name – identifies the genus to which the species belongs, whereas the second part – the specific name or specific epithet – distinguishes the species within the genus. For example, modern humans belong to the genus Homo and within this genus to the species Homo sapiens. Tyrannosaurus rex is likely the most widely known binomial.BOOK, Busby, Arthur III, etal, A Guide to Rocks and Fossils, 103, 1997, The formal introduction of this system of naming species is credited to Carl Linnaeus, effectively beginning with his work Species Plantarum in 1753. But as early as 1622, Gaspard Bauhin introduced in his book Pinax theatri botanici (English, Illustrated exposition of plants) containing many names of genera that were later adopted by Linnaeus.WEB, Bauhin, Gaspard, Pinax theatri botanici,edb.kulib.kyoto-u.ac.jp/exhibit-e/b07/b07cont.html, Kyoto University Library, 19 June 2016, 17 October 2016,edb.kulib.kyoto-u.ac.jp/exhibit-e/b07/b07cont.html," title="web.archive.org/web/20161017200341edb.kulib.kyoto-u.ac.jp/exhibit-e/b07/b07cont.html,">web.archive.org/web/20161017200341edb.kulib.kyoto-u.ac.jp/exhibit-e/b07/b07cont.html, live, The application of binomial nomenclature is now governed by various internationally agreed codes of rules, of which the two most important are the International Code of Zoological Nomenclature (ICZN) for animals and the International Code of Nomenclature for algae, fungi, and plants (ICNafp or ICN). Although the general principles underlying binomial nomenclature are common to these two codes, there are some differences in the terminology they use and their particular rules.In modern usage, the first letter of the generic name is always capitalized in writing, while that of the specific epithet is not, even when derived from a proper noun such as the name of a person or place. Similarly, both parts are italicized in normal text (or underlined in handwriting). Thus the binomial name of the annual phlox (named after botanist Thomas Drummond) is now written as Phlox drummondii. Often, after a species name is introduced in a text, the generic name is abbreviated to the first letter in subsequent mentions (e.g., P. drummondii).In scientific works, the authority for a binomial name is usually given, at least when it is first mentioned, and the year of publication may be specified.
  • In zoology
    • Patella vulgata Linnaeus, 1758”. The name “Linnaeus” tells the reader who published the name and description for this species; 1758 is the year the name and original description were published (in this case, in the 10th edition of the book Systema Naturae).
    • Passer domesticus (Linnaeus, 1758)”. The original name given by Linnaeus was Fringilla domestica; the parentheses indicate that the species is now placed in a different genus. The ICZN does not require that the name of the person who changed the genus be given, nor the date on which the change was made, although nomenclatorial catalogs usually include such information.
  • In botany
    • Amaranthus retroflexus L.“ â€“ “L.” is the standard abbreviation used for “Linnaeus”.
    • Hyacinthoides italica (L.) Rothm.“ â€“ Linnaeus first named this bluebell species Scilla italica; Rothmaler transferred it to the genus Hyacinthoides; the ICNafp does not require that the dates of either publication be specified.

Etymology

The name is composed of two word-forming elements: {{wikt-lang|la|bi-}} (Latin prefix meaning ‘two’) and {{wikt-lang|la|nomial}} (the adjective form of , Latin for ‘name’). In Medieval Latin, the related word was used to signify one term in a binomial expression in mathematics.WEB,www.cnrtl.fr/definition/bin%C3%B4me, binôme, Le Trésor de la langue française informatisé, Centre National de Ressources Textuelles et Lexicales,www.cnrtl.fr/definition/bin%C3%B4me," title="web.archive.org/web/20170606122907www.cnrtl.fr/definition/bin%C3%B4me,">web.archive.org/web/20170606122907www.cnrtl.fr/definition/bin%C3%B4me, 6 June 2017, In fact, the Latin word may validly refer to either of the epithets in the binomial name, which can equally be referred to as a {{wikt-lang|la|binomen}} (pl. {{wikt-lang|la|binomina}}).{{better source needed|date=December 2023|reason=Wikitionary fails , like WP itself does. This must be in other, independent dictionaries.}}

History

{{see also|Biological classification#Early systems||label 1=Biological classification: Early systems}}File:Carl von Linné.jpg|thumb|Carl LinnaeusCarl LinnaeusPrior to the adoption of the modern binomial system of naming species, a scientific name consisted of a generic name combined with a specific name that was from one to several words long. Together they formed a system of polynomial nomenclature.BOOK, Reddy, S. M., University botany: Angiosperms, plant embryology and plant physiology,books.google.com/books?id=SuQJeFkYoRIC&pg=PA34, 2007, New Age International, 978-81-224-1547-6, 34, 13 November 2019, 2 February 2023,web.archive.org/web/20230202160908/https://books.google.com/books?id=SuQJeFkYoRIC&pg=PA34, live, These names had two separate functions. First, to designate or label the species, and second, to be a diagnosis or description; however, these two goals were eventually found to be incompatible.BOOK, Blunt, Wilfrid, Linnaeus: The compleat naturalist,books.google.com/books?id=FRH_EMhQYhYC&pg=PA266, 2004, Frances Lincoln Ltd, 978-0-7112-2362-2, 266, In a simple genus, containing only two species, it was easy to tell them apart with a one-word genus and a one-word specific name; but as more species were discovered, the names necessarily became longer and unwieldy, for instance, Plantago foliis ovato-lanceolatus pubescentibus, spica cylindrica, scapo tereti (“plantain with pubescent ovate-lanceolate leaves, a cylindric spike and a terete scape“), which we know today as Plantago media.{{cn|date=November 2023}}Such “polynomial names” may sometimes look like binomials, but are significantly different. For example, Gerard’s herbal (as amended by Johnson) describes various kinds of spiderwort: “The first is called Phalangium ramosum, Branched Spiderwort; the second, Phalangium non ramosum, Unbranched Spiderwort. The other ... is aptly termed Phalangium Ephemerum Virginianum, Soon-Fading Spiderwort of Virginia”.BOOK, John, Gerard, Thomas, Johnson, 1636, The Herball, or, Generall Historie of Plantes /gathered by John Gerarde of London, Master in Chirurgerie; very much enlarged and amended by Thomas Johnson, Citizen and Apothecarye of London, Adam Islip, Joice Norton and Richard Whitakers and the Biodiversity Heritage Library,www.biodiversitylibrary.org/item/33580#page/88/mode/1up, 14 January 2018, 11 October 2017,web.archive.org/web/20171011132117/https://www.biodiversitylibrary.org/item/33580#page/88/mode/1up, live, The Latin phrases are short descriptions, rather than identifying labels.The Bauhins, in particular Caspar Bauhin (1560–1624), took some important steps towards the binomial system, by pruning the Latin descriptions, in many cases to two words.BOOK, Johnson, A. T., Smith, H. A., Stockdale, A. P., 2019, Plant Names Simplified: Their Pronunciation Derivation & Meaning, Sheffield, Yorkshire, 5M Publishing, 978-1-910455-06-7, , p. v The adoption by biologists of a system of strictly binomial nomenclature is due to Swedish botanist and physician Carl Linnaeus (1707–1778). {{anchor|trivial name}} It was in Linnaeus’s 1753 Species Plantarum that he began consistently using a one-word trivial name () after a generic name (genus name) in a system of binomial nomenclature.BOOK, Polaszek, Andrew, Systema naturae 250: The Linnaean ark,books.google.com/books?id=ReWP31_IJSIC&pg=PA189, 2009, CRC Press, 978-1-4200-9501-2, 189, Trivial names had already appeared in his Critica Botanica (1737) and Philosophia Botanica (1751). This trivial name is what is now known as a specific epithet (ICNafp) or specific name (ICZN). The Bauhins’ genus names were retained in many of these, but the descriptive part was reduced to a single word.Linnaeus’s trivial names introduced an important new idea, namely that the function of a name could simply be to give a species a unique label. This meant that the name no longer needs to be descriptive; for example, both parts could be derived from the names of people. Thus Gerard’s Phalangium ephemerum virginianum became Tradescantia virginiana, where the genus name honoured John Tradescant the Younger,{{NoteTag|Some sources say that both John Tradescant the Younger and his father, John Tradescant the Elder, were intended by Linnaeus.}} an English botanist and gardener.{{Harvnb|Hyam|Pankhurst|1995|p=502}} A bird in the parrot family was named Psittacus alexandri, meaning “Alexander’s parrot”, after Alexander the Great, whose armies introduced eastern parakeets to Greece.BOOK, Jobling, James A., 2010, Helm Dictionary of Scientific Bird Names, London, Christopher Helm, 978-1-4081-2501-4, Linnaeus’s trivial names were much easier to remember and use than the parallel polynomial names and eventually replaced them.WEB, Knapp, Sandra, Sandra Knapp, What’s in a name? A history of taxonomy: Linnaeus and the birth of modern taxonomy, Natural History Museum, London, NHM.ac.uk,www.nhm.ac.uk/nature-online/science-of-natural-history/taxonomy-systematics/history-taxonomy/session1/index.html, 17 June 2011,www.nhm.ac.uk/nature-online/science-of-natural-history/taxonomy-systematics/history-taxonomy/session1/index.html," title="web.archive.org/web/20141018181831www.nhm.ac.uk/nature-online/science-of-natural-history/taxonomy-systematics/history-taxonomy/session1/index.html,">web.archive.org/web/20141018181831www.nhm.ac.uk/nature-online/science-of-natural-history/taxonomy-systematics/history-taxonomy/session1/index.html, 18 October 2014,

Value

File:E coli at 10000x, original.jpg|thumb|The bacterium Escherichia coliEscherichia coliThe value of the binomial nomenclature system derives primarily from its economy, its widespread use, and the uniqueness and stability of names that the Codes of Zoological and Botanical, Bacterial and Viral Nomenclature provide:
  • Economy. Compared to the polynomial system which it replaced, a binomial name is shorter and easier to remember. It corresponds to the widespread system of family name plus given name(s) used to name people in many cultures.
  • Widespread use. The binomial system of nomenclature is governed by international codes and is used by biologists worldwide.BOOK, Van Dyke, Fred, 2008, Contemporary Issues of the Species Concept, Conservation biology: foundations, concepts, applications, 86, Springer, 978-1-4020-6890-4,books.google.com/books?id=Evh1UD3ZYWcC&pg=PA86, 20 June 2011, A few binomials have also entered common speech, such as Homo sapiens, E. coli, Boa constrictor, Tyrannosaurus rex, and Aloe vera.
  • Uniqueness. Provided that taxonomists agree as to the limits of a species, it can have only one name that is correct under the appropriate nomenclature code, generally the earliest published if two or more names are accidentally assigned to a species.BOOK, Peter J., Russell, Stephen L., Wolfe, Paul E., Hertz, Cecie, Starr, The Linnaean System of Taxonomy, 2, Cengage Learning, Species Concepts and Speciation, 2007, 493, 978-0-495-01033-3,books.google.com/books?id=7F9UlGYGr0sC&pg=PA493, However, establishing that two names actually refer to the same species and then determining which has priority can be difficult, particularly if the species was named by biologists from different countries. Therefore, a species may have more than one regularly used name; all but one of these names are “synonyms”.JOURNAL, 2007, General Principles of Taxonomy, Competition Science Vision, 10, 114, 764–767,books.google.com/books?id=0-cDAAAAMBAJ, 20 June 2011, Darpan, Pratiyogita, Furthermore, within zoology or botany, each species name applies to only one species. If a name is used more than once, it is called a homonym.
File:Petirrojo (Erithacus rubecula superbus ) (6178023045).jpg|thumb|Erithacus rubecula superbus, the Tenerife robin or petirrojo]]

Problems

Binomial nomenclature for species has the effect that when a species is moved from one genus to another, sometimes the specific name or epithet must be changed as well. This may happen because the specific name is already used in the new genus, or to agree in gender with the new genus if the specific epithet is an adjective modifying the genus name. Some biologists have argued for the combination of the genus name and specific epithet into a single unambiguous name, or for the use of uninomials (as used in nomenclature of ranks above species).JOURNAL, Michener, Charles D., 1964, The possible use of uninominal nomenclature to increase the stability of names in biology, Systematic Zoology, 13, 4, 182–190, 10.2307/2411777, 2411777, JOURNAL, Cantino, P. D., H. D., Bryant, K., de Queiroz, M. J., Donoghue, T., Eriksson, D. M., Hillis, M. S. Y., Lee, 1999,www.botany.wisc.edu/courses/botany_940/05PhyloCode/papers/Cantino%2699b.pdf,www.botany.wisc.edu/courses/botany_940/05PhyloCode/papers/Cantino%2699b.pdf," title="ghostarchive.org/archive/20221009www.botany.wisc.edu/courses/botany_940/05PhyloCode/papers/Cantino%2699b.pdf,">ghostarchive.org/archive/20221009www.botany.wisc.edu/courses/botany_940/05PhyloCode/papers/Cantino%2699b.pdf, 9 October 2022, live, Species names in phylogenetic nomenclature, Systematic Biology, 48, 4, 790–807, 10.1080/106351599260012, 12066299, free, Because genus names are unique only within a nomenclature code, it is possible for homonyms (two or more species sharing the same genus name) to happen, and even the same binomial if they occur in different kingdoms. At least 1,258 instances of genus name duplication occur (mainly between zoology and botany).WEB, HemiHomonym Database,herba.msu.ru/shipunov/os/homonyms/index.php, herba.msu.ru, 17 November 2022,herba.msu.ru/shipunov/os/homonyms/index.php," title="web.archive.org/web/20150512143402herba.msu.ru/shipunov/os/homonyms/index.php,">web.archive.org/web/20150512143402herba.msu.ru/shipunov/os/homonyms/index.php, 12 May 2015, JOURNAL, Shipunov, Alexey, The problem of hemihomonyms and the on-line hemihomonyms database (HHDB), Bionomina, 16 January 2013, 4, 1, 65–72, 10.11646/bionomina.4.1.3, free,

Relationship to classification and taxonomy

Nomenclature (including binomial nomenclature) is not the same as classification, although the two are related. Classification is the ordering of items into groups based on similarities or differences; in biological classification, species are one of the kinds of item to be classified.BOOK, Simpson, Michael G., 2006, Plant Systematics, London, Elsevier Academic Press, 978-0-12-644460-5, , p. 552 In principle, the names given to species could be completely independent of their classification. This is not the case for binomial names, since the first part of a binomial is the name of the genus into which the species is placed. Above the rank of genus, binomial nomenclature and classification are partly independent; for example, a species retains its binomial name if it is moved from one family to another or from one order to another, unless it better fits a different genus in the same or different family, or it is split from its old genus and placed in a newly created genus. The independence is only partial since the names of families and other higher taxa are usually based on genera.{{Citation needed|date=June 2011}}Taxonomy includes both nomenclature and classification. Its first stages (sometimes called “alpha taxonomy“) are concerned with finding, describing and naming species of living or fossil organisms.{{Citation |last=Fortey |first=Richard |author-link=Richard Fortey |date=2008 |title=Dry Store Room No. 1: The Secret Life of the Natural History Museum |location=London |publisher=Harper Perennial |isbn=978-0-00-720989-7}} Binomial nomenclature is thus an important part of taxonomy as it is the system by which species are named. Taxonomists are also concerned with classification, including its principles, procedures and rules.BOOK, Davis, Peter H., Peter Hadland Davis, Heywood, Vernon H., Vernon Heywood, 1965, Principles of Angiosperm Taxonomy, Edinburgh, Oliver & Boyd, 8,

Derivation of binomial names

{{see also|List of Latin and Greek words commonly used in systematic names}}A complete binomial name is always treated grammatically as if it were a phrase in the Latin language (hence the common use of the term “Latin name” for a binomial name). However, the two parts of a binomial name can each be derived from a number of sources, of which Latin is only one. These include: The first part of the name, which identifies the genus, must be a word which can be treated as a Latin singular noun in the nominative case. It must be unique within the purview of each nomenclatural code, but can be repeated between them. Thus Huia recurvata is an extinct species of plant, found as fossils in Yunnan, China,JOURNAL, Geng, Bao-Yin, 1985, Huia recurvata â€“ A New Plant from Lower Devonian of Southeastern Yunnan China, Acta Botanica Sinica, 27, 4, 419–426,en.cnki.com.cn/Article_en/CJFDTOTAL-ZWXB198504013.htm, 7 February 2011, zh, en, 7 July 2011,en.cnki.com.cn/Article_en/CJFDTOTAL-ZWXB198504013.htm," title="web.archive.org/web/20110707014652en.cnki.com.cn/Article_en/CJFDTOTAL-ZWXB198504013.htm,">web.archive.org/web/20110707014652en.cnki.com.cn/Article_en/CJFDTOTAL-ZWXB198504013.htm, live, whereas Huia masonii is a species of frog found in Java, Indonesia.WEB, Iskandar, D., Mumpuni, D., 2004, IUCN Red List of Threatened Species: Huia masonii, IUCNRedList.org, International Union for Conservation of Nature,iucnredlist.org/details/58303/0, 19 June 2011, 26 October 2012,www.iucnredlist.org/details/58303/0," title="web.archive.org/web/20121026102204www.iucnredlist.org/details/58303/0,">web.archive.org/web/20121026102204www.iucnredlist.org/details/58303/0, live, The second part of the name, which identifies the species within the genus, is also treated grammatically as a Latin word. It can have one of a number of forms:
  • The second part of a binomial may be an adjective. The adjective must agree with the genus name in gender. Latin has three genders, masculine, feminine and neuter, shown by varying endings to nouns and adjectives. The house sparrow has the binomial name Passer domesticus. Here domesticus (“domestic“) simply means “associated with the house”. The sacred bamboo is Nandina domestica{{Harvnb|Hyam|Pankhurst|1995|p=334}} rather than Nandina domesticus, since Nandina is feminine whereas Passer is masculine. The tropical fruit langsat is a product of the plant Lansium parasiticum, since Lansium is neuter. Some common endings for Latin adjectives in the three genders (masculine, feminine, neuter) are -us, -a, -um (as in the previous example of domesticus); -is, -is, -e (e.g., tristis, meaning “sad“); and -or -or -us (e.g., minor, meaning “smaller“). For further information, see (Latin declensionAdjectives|Latin declension: Adjectives).
  • The second part of a binomial may be a noun in the nominative case. An example is the binomial name of the lion, which is Panthera leo. Grammatically the noun is said to be in apposition to the genus name and the two nouns do not have to agree in gender; in this case, Panthera is feminine and leo is masculine.
File:Talauma hodgsonii.jpg|thumb|Magnolia hodgsoniiMagnolia hodgsonii
  • The second part of a binomial may be a noun in the genitive (possessive) case. The genitive case is constructed in a number of ways in Latin, depending on the declension of the noun. Common endings for masculine and neuter nouns are -ii or -i in the singular and -orum in the plural, and for feminine nouns -ae in the singular and -arum in the plural. The noun may be part of a person’s name, often the surname, as in the Tibetan antelope (Pantholops hodgsonii), the shrub Magnolia hodgsonii, or the olive-backed pipit (Anthus hodgsoni). The meaning is “of the person named”, so Magnolia hodgsonii means “Hodgson’s magnolia”. The -ii or -i endings show that in each case Hodgson was a man (not the same one); had Hodgson been a woman, hodgsonae would have been used. The person commemorated in the binomial name is not usually (if ever) the person who created the name; for example, Anthus hodgsoni was named by Charles Wallace Richmond, in honour of Hodgson. Rather than a person, the noun may be related to a place, as with Latimeria chalumnae, meaning “of the Chalumna River”. Another use of genitive nouns is in, for example, the name of the bacterium Escherichia coli, where coli means “of the colon”. This formation is common in parasites, as in Xenos vesparum, where vesparum means “of the wasps”, since Xenos vesparum is a parasite of wasps.
Whereas the first part of a binomial name must be unique within the purview of each nomenclatural code, the second part is quite commonly used in two or more genera (as is shown by examples of hodgsonii above). The full binomial name must be unique within each code.

Codes

From the early 19th century onwards it became ever more apparent that a body of rules was necessary to govern scientific names. In the course of time these became nomenclature codes. The International Code of Zoological Nomenclature (ICZN) governs the naming of animals,{{Harvnb|International Commission on Zoological Nomenclature|1999}} the International Code of Nomenclature for algae, fungi, and plants (ICNafp) that of plants (including cyanobacteria), and the International Code of Nomenclature of Bacteria (ICNB) that of bacteria (including Archaea). Virus names are governed by the International Committee on Taxonomy of Viruses (ICTV), a taxonomic code, which determines taxa as well as names. These codes differ in certain ways, e.g.: ! Code !! Full name !! First part !! Second part
| specific name
| specific epithet
Unifying the different codes into a single code, the ”BioCode”, has been suggested{{By whom|date=November 2022}}, although implementation is not in sight. (There is also a published code for a different system of biotic nomenclature, which does not use ranks above species, but instead names clades. This is called PhyloCode.)

Differences in handling personal names

As noted above, there are some differences between the codes in how binomials can be formed; for example the ICZN allows both parts to be the same, while the ICNafp does not. Another difference is in how personal names are used in forming specific names or epithets. The ICNafp sets out precise rules by which a personal name is to be converted to a specific epithet. In particular, names ending in a consonant (but not “er“) are treated as first being converted into Latin by adding “-ius” (for a man) or “-ia” (for a woman), and then being made genitive (i.e. meaning “of that person or persons“). This produces specific epithets like lecardii for Lecard (male), wilsoniae for Wilson (female), and brauniarum for the Braun sisters.McNeill et al. 2012, Recommendation 60C By contrast, the ICZN does not require the intermediate creation of a Latin form of a personal name, allowing the genitive ending to be added directly to the personal name.{{Harvnb|International Commission on Zoological Nomenclature|1999}}, Chap. 7, Article 3.1.2 This explains the difference between the names of the plant Magnolia hodgsonii and the bird Anthus hodgsoni. Furthermore, the ICNafp requires names not published in the form required by the code to be corrected to conform to it,McNeill et al. 2012, Article 60.12 whereas the ICZN is more protective of the form used by the original author.{{Harvnb|International Commission on Zoological Nomenclature|1999}}, Chap. 7, Article 32.3

Writing binomial names

By tradition, the binomial names of species are usually typeset in italics; for example, Homo sapiens.JOURNAL, How to Write Scientific Names of Organisms, Competition Science Vision,www.journal.au.edu/au_techno/2001/oct2001/howto.pdf, 20 June 2011, 4 July 2011,www.journal.au.edu/au_techno/2001/oct2001/howto.pdf," title="web.archive.org/web/20110704203742www.journal.au.edu/au_techno/2001/oct2001/howto.pdf,">web.archive.org/web/20110704203742www.journal.au.edu/au_techno/2001/oct2001/howto.pdf, live, Generally, the binomial should be printed in a font style different from that used in the normal text; for example, ”Several more Homo sapiens fossils were discovered.” When handwritten, a binomial name should be underlined; for example, Homo sapiens.WEB,www.cdtl.nus.edu.sg/success/sl43.htm, Understanding and Learning Scientific Names of Species, Tan, Hugh T.W., Tan, Kai-xin, Successful Learning, Center for Development of Teaching and Learning, National University of Singapore, CDTL.NUS.edu.sg, 20 June 2011, 21 July 2011,www.cdtl.nus.edu.sg/success/sl43.htm," title="web.archive.org/web/20110721013443www.cdtl.nus.edu.sg/success/sl43.htm,">web.archive.org/web/20110721013443www.cdtl.nus.edu.sg/success/sl43.htm, live, The first part of the binomial, the genus name, is always written with an initial capital letter. Older sources, particularly botanical works published before the 1950s, used a different convention: if the second part of the name was derived from a proper noun, e.g., the name of a person or place, a capital letter was used. Thus, the modern form Berberis darwinii was written as Berberis Darwinii. A capital was also used when the name is formed by two nouns in apposition, e.g., Panthera Leo or Centaurea Cyanus.{{sfn|Johnson|Smith|1972|p=23}}{{NoteTag|The modern notation was resisted by some, partly because writing names like Centaurea cyanus can suggest that cyanus is an adjective which should agree with Centaurea, i.e. that the name should be Centaurea cyana, whereas Cyanus is derived from the Greek name for the cornflower.BOOK, Gilbert-Carter, H., 1955, Glossary of the British Flora, 2nd, Cambridge University Press, 559413416, xix, }} In current usage, the second part is never written with an initial capital.BOOK, Silyn-Roberts, Heather, 2000, Writing for Science and Engineering: Papers, Presentations and Reports, Oxford; Boston, Butterworth-Heinemann, 978-0-7506-4636-9, 198, McNeill et al. 2012, Recommendation 60FWhen used with a common name, the scientific name often follows in parentheses, although this varies with publication.BOOK,www.gse.mq.edu.au/resources/guide/format.htm, Writing Guide: Language, Words and Format, Sydney, NSW, Macquarie University, 29 June 2011,www.gse.mq.edu.au/resources/guide/format.htm," title="web.archive.org/web/20110629140234www.gse.mq.edu.au/resources/guide/format.htm,">web.archive.org/web/20110629140234www.gse.mq.edu.au/resources/guide/format.htm, For example, “The house sparrow (Passer domesticus) is decreasing in Europe.“The binomial name should generally be written in full. The exception to this is when several species from the same genus are being listed or discussed in the same paper or report, or the same species is mentioned repeatedly; in which case the genus is written in full when it is first used, but may then be abbreviated to an initial (and a period/full stop).BOOK, Merriam-Webster’s Collegiate Dictionary, Merriam-Webster, Linnaean Nomenclature of Plants, Animals, & Bacteria, 2003, 22a â€“ 23a, 978-0-87779-809-5,books.google.com/books?id=TAnheeIPcAEC&pg=PP28, registration,archive.org/details/merriamwebstersc00merr_6/page/22, For example, a list of members of the genus Canis might be written as ”Canis lupus, C. aureus, C. simensis”. In rare cases, this abbreviated form has spread to more general use; for example, the bacterium Escherichia coli is often referred to as just E. coli, and Tyrannosaurus rex is perhaps even better known simply as T. rex, these two both often appearing in this form in popular writing even where the full genus name has not already been given.The abbreviation “sp.” is used when the actual specific name cannot or need not be specified. The abbreviation “spp.” (plural) indicates “several species”. These abbreviations are not italicised (or underlined).BOOK, Nair, P. K. Ramachandran, P. K. Ramachandran Nair, Nair, Vimala D., 2014, Scientific Writing and Communication in Agriculture and Natural Resources,books.google.com/books?id=Uxm4BAAAQBAJ&pg=PA39, Springer Science+Business Media, Springer International Publishing, 39, 10.1007/978-3-319-03101-9, 978-3-319-03101-9, 11811479, 2013953625, 881314963, 20 March 2023, 28 March 2023,web.archive.org/web/20230328005811/https://books.google.com/books?id=Uxm4BAAAQBAJ&pg=PA39, live, WEB,www.hort.purdue.edu/hort/courses/hort217/Nomenclature/description.htm,www.hort.purdue.edu/hort/courses/hort217/Nomenclature/description.htm," title="archive.today/20121211140110www.hort.purdue.edu/hort/courses/hort217/Nomenclature/description.htm,">archive.today/20121211140110www.hort.purdue.edu/hort/courses/hort217/Nomenclature/description.htm, 11 December 2012, Plant Nomenclature, Matthew A., Jenks, Department of Horticulture and Landscape Architecture, Purdue University, Hort.Purdue.edu, 20 June 2011, For example: ”Canis sp.” means “an unspecified species of the genus Canis”, while ”Canis spp.” means “two or more species of the genus Canis”. (These abbreviations should not be confused with the abbreviations “ssp.” (zoology) or “subsp.” (botany), plurals “sspp.” or “subspp.”, referring to one or more subspecies. See trinomen (zoology) and infraspecific name.)The abbreviation “cf.” (i.e., confer in Latin) is used to compare individuals/taxa with known/described species. Conventions for use of the “cf.” qualifier vary.WEB,www.peripatus.gen.nz/biology/priTaxonomy.html, Taxonomy â€“ A Primer, Chris, Clowes, Peripatus.Gen.nz, 20 June 2011, 7 August 2011,web.archive.org/web/20110807091803/https://www.peripatus.gen.nz/biology/priTaxonomy.html, In paleontology, it is typically used when the identification is not confirmed.JOURNAL, Bengtson, P., 1988, Open nomenclature, Palaeontology, 31, 1, 223–227,www.palass-pubs.org/palaeontology/pdf/Vol31/Pages%20223-227.pdf,www.palass-pubs.org/palaeontology/pdf/Vol31/Pages%20223-227.pdf," title="web.archive.org/web/20141006071437www.palass-pubs.org/palaeontology/pdf/Vol31/Pages%20223-227.pdf,">web.archive.org/web/20141006071437www.palass-pubs.org/palaeontology/pdf/Vol31/Pages%20223-227.pdf, 6 October 2014, 22 August 2014, For example, ”Corvus cf. nasicus” was used to indicate “a fossil bird similar to the Cuban crow but not certainly identified as this species”.JOURNAL, Orihuela, J., 2013,birdscaribbean.org/jco/index.php/jco/article/view/32, Fossil Cuban crow Corvus cf. nasicus from a late Quaternary cave deposit in northern Matanzas, Cuba, Journal of Caribbean Ornithology, 26, 12–16, 22 August 2014, 27 January 2018,birdscaribbean.org/jco/index.php/jco/article/view/32," title="web.archive.org/web/20180127004555birdscaribbean.org/jco/index.php/jco/article/view/32,">web.archive.org/web/20180127004555birdscaribbean.org/jco/index.php/jco/article/view/32, live, In molecular systematics papers, “cf.” may be used to indicate one or more undescribed species assumed to be related to a described species. For example, in a paper describing the phylogeny of small benthic freshwater fish called darters, five undescribed putative species (Ozark, Sheltowee, Wildcat, Ihiyo, and Mamequit darters), notable for brightly colored nuptial males with distinctive color patterns,BOOK, Page, L. M., Burr, B. M., 1991, Peterson field guide to freshwater fishes: North America north of Mexico,archive.org/details/fieldguidetofres00lawr, registration, Houghton Mifflin Harcourt, 978-0-547-24206-4, plate 52, were referred to as ”Etheostoma cf. spectabile” because they had been viewed as related to, but distinct from, Etheostoma spectabile (orangethroat darter).JOURNAL, Near, T. J., Bossu, C. M., Bradburd, G. S., Carlson, R. L., Harrington, R. C., Hollingsworth, P. R., Keck, B. P., Etnier, D. A., 2011, Phylogeny and temporal diversification of darters (Percidae: Etheostomatinae), Systematic Biology, 60, 5, 565–595, 10.1093/sysbio/syr052, free, 21775340, This view was supported to varying degrees by DNA analysis. The somewhat informal use of taxa names with qualifying abbreviations is referred to as open nomenclature and it is not subject to strict usage codes.In some contexts, the dagger symbol (“†“) may be used before or after the binomial name to indicate that the species is extinct.{{anchor|Authority}}

Authority

In scholarly texts, at least the first or main use of the binomial name is usually followed by the “authority“ â€“ a way of designating the scientist(s) who first published the name. The authority is written in slightly different ways in zoology and botany. For names governed by the ICZN the surname is usually written in full together with the date (normally only the year) of publication. One example of author citation of scientific name is: “Amabela Möschler, 1880.“{{NoteTag|Here Amabela is the name of the genus. It is written in italic form. Followed by the last name of the scientist who discovered it (Heinrich Benno Möschler), a comma, and the year when it was published.}} The ICZN recommends that the “original author and date of a name should be cited at least once in each work dealing with the taxon denoted by that name.“{{sfn|International Commission on Zoological Nomenclature |1999|loc=Recommendation 51a}} For names governed by the ICNafp the name is generally reduced to a standard abbreviation and the date omitted. The International Plant Names Index maintains an approved list of botanical author abbreviations. Historically, abbreviations were used in zoology too.When the original name is changed, e.g., the species is moved to a different genus, both codes use parentheses around the original authority; the ICNafp also requires the person who made the change to be given. In the ICNafp, the original name is then called the basionym. Some examples:
  • (Plant) Amaranthus retroflexus L. â€“ “L.” is the standard abbreviation for “Linnaeus”; the absence of parentheses shows that this is his original name.
  • (Plant) Hyacinthoides italica (L.) Rothm. â€“ Linnaeus first named the Italian bluebell Scilla italica; that is the basionym. Rothmaler later transferred it to the genus Hyacinthoides.
  • (Animal) Passer domesticus (Linnaeus, 1758) â€“ the original name given by Linnaeus was Fringilla domestica; unlike the ICNafp, the ICZN does not require the name of the person who changed the genus (Mathurin Jacques BrissonBOOK, Summers-Smith, J. Denis, The Sparrows, T. & A. D. Poyser, 1988, Calton, Staffordshire, England, 978-0-85661-048-6, illustrated by Robert Gillmor, registration,archive.org/details/sparrowsstudyofg0000summ, 114–115, ) to be given.

Other ranks

{{unreferenced section|date=June 2011}}Binomial nomenclature, as described here, is a system for naming species. Implicitly, it includes a system for naming genera, since the first part of the name of the species is a genus name. In a classification system based on ranks, there are also ways of naming ranks above the level of genus and below the level of species. Ranks above genus (e.g., family, order, class) receive one-part names, which are conventionally not written in italics. Thus, the house sparrow, Passer domesticus, belongs to the family Passeridae. Family names are normally based on genus names, although the endings used differ between zoology and botany.Ranks below species receive three-part names, conventionally written in italics like the names of species. There are significant differences between the ICZN and the ICNafp. In zoology, the only formal rank below species is subspecies and the name is written simply as three parts (a trinomen). Thus, one of the subspecies of the olive-backed pipit is Anthus hodgsoni berezowskii. Informally, in some circumstances, a form may be appended. For example Harmonia axyridis f. spectabilis is the harlequin ladybird in its black or melanic forms having four large orange or red spots. In botany, there are many ranks below species and although the name itself is written in three parts, a “connecting term” (not part of the name) is needed to show the rank. Thus, the American black elder is Sambucus nigra subsp. canadensis; the white-flowered form of the ivy-leaved cyclamen is Cyclamen hederifolium f. albiflorum.

See also

Notes

{{NoteFoot}}

References

{{Reflist}}

Bibliography

  • BOOK, Hyam, R., Pankhurst, R. J., 1995, Plants and their names: A concise dictionary, Oxford, Oxford University Press, 978-0-19-866189-4,
  • BOOK, Johnson, A.T., Smith, H. A., 1972, Plant Names Simplified: Their Pronunciation Derivation & Meaning, Buckenhill, Herefordshire, Landsmans Bookshop, 978-0-900513-04-6,
  • BOOK, International Commission on Zoological Nomenclature, 1999, International Code of Zoological Nomenclature online, 4th, The International Trust for Zoological Nomenclature, 978-0-85301-006-7,www.nhm.ac.uk/hosted-sites/iczn/code/, 20 June 2011,
  • BOOK, McNeill, J., Barrie, F. R., Buck, W. R., Demoulin, V., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Marhold, K., Prado, J., Prud’homme Van Reine, W. F., Smith, G. F., Wiersema, J. H., Turland, N. J., 2012, Regnum Vegetabile 154, International Code of Nomenclature for algae, fungi, and plants (Melbourne Code) adopted by the Eighteenth International Botanical Congress Melbourne, Australia, July 2011, Königstein, Germany, Koeltz Scientific Books, 978-3-87429-425-6, ICN2012,www.iapt-taxon.org/nomen/main.php?page=title, deviated, 7 May 2017,www.iapt-taxon.org/nomen/main.php?page=title," title="web.archive.org/web/20170507070323www.iapt-taxon.org/nomen/main.php?page=title,">web.archive.org/web/20170507070323www.iapt-taxon.org/nomen/main.php?page=title,

Further reading

External links

{{Commons category|Biological nomenclature}} {{Carl Linnaeus}}{{Authority control}}{{Interwiki extra|qid=Q10753560}}

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