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ecosystem
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{{pp-vandalism|small=yes}}{{short description|A community of living organisms together with the nonliving components of their environment}}{{about|natural ecosystems|the term used in man-made systems|Digital ecosystem|the system of classifying ecologically homogeneous land units|Biome}}{{redirect|Biosystem|the journal|BioSystems}}{{Multiple image
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| image1 = Blue Linckia Starfish.JPG
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| alt1 = Coral reefs are a highly productive marine ecosystem.|image2=HohRiverTrail 7322.jpg|width2=180
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| footer = Left: Coral reef ecosystems are highly productive marine systems.JOURNAL, Hatcher, Bruce Gordon, 1990, Coral reef primary productivity. A hierarchy of pattern and process, Trends in Ecology and Evolution, 5, 5, 149–155, 10.1016/0169-5347(90)90221-X, 21232343, , right: Temperate rainforest on the Olympic Peninsula in Washington state.
}}An ecosystem is a community of living organisms in conjunction with the nonliving components of their environment, interacting as a system.Tansley (1934); Molles (1999), p. 482; Chapin et al. (2002), p. 380; Schulze et al. (2005); p. 400; Gurevitch et al. (2006), p. 522; Smith & Smith 2012, p. G-5 These biotic and abiotic components are linked together through nutrient cycles and energy flows.BOOK, Fundamentals of Ecology, Odum, Eugene P, 1971, Saunders, 978-0-534-42066-6, third, New York, Energy enters the system through photosynthesis and is incorporated into plant tissue. By feeding on plants and on one-another, animals play an important role in the movement of matter and energy through the system. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter, decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to a form that can be readily used by plants and other microbes.Chapin et al. (2002), p. 10Ecosystems are controlled by external and internal factors. External factors such as climate, parent material which forms the soil and topography, control the overall structure of an ecosystem but are not themselves influenced by the ecosystem.Chapin et al. (2002), pp. 11–13 Unlike external factors, internal factors are controlled, for example, decomposition, root competition, shading, disturbance, succession, and the types of species present. Ecosystems are dynamic entities—they are subject to periodic disturbances and are in the process of recovering from some past disturbance.Chapin et al. (2002), pp. 281–304 Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present. Internal factors not only control ecosystem processes but are also controlled by them and are often subject to feedback loops.Resource inputs are generally controlled by external processes like climate and parent material. Resource availability within the ecosystem is controlled by internal factors like decomposition, root competition or shading. Although humans operate within ecosystems, their cumulative effects are large enough to influence external factors like climate.Biodiversity affects ecosystem functioning, as do the processes of disturbance and succession. Ecosystems provide a variety of goods and services upon which people depend.

History

The term ecosystem was first used in 1935 in a publication by British ecologist Arthur Tansley.The term ecosystem was actually coined by Arthur Roy Clapham, who came up with the word at Tansley's request (Willis 1997).Willis (1997) Tansley devised the concept to draw attention to the importance of transfers of materials between organisms and their environment.Chapin et al. (2002), pp. 7–11) He later refined the term, describing it as "The whole system, ... including not only the organism-complex, but also the whole complex of physical factors forming what we call the environment".Tansley (1935) Tansley regarded ecosystems not simply as natural units, but as "mental isolates". Tansley later defined the spatial extent of ecosystems using the term ecotope.BOOK,weblink The British Islands and Their Vegetation, Tansley, A.G., Cambridge University Press, 1939, Arthur Tansley, G. Evelyn Hutchinson, a limnologist who was a contemporary of Tansley's, combined Charles Elton's ideas about trophic ecology with those of Russian geochemist Vladimir Vernadsky. As a result, he suggested that mineral nutrient availability in a lake limited algal production. This would, in turn, limit the abundance of animals that feed on algae. Raymond Lindeman took these ideas further to suggest that the flow of energy through a lake was the primary driver of the ecosystem. Hutchinson's students, brothers Howard T. Odum and Eugene P. Odum, further developed a "systems approach" to the study of ecosystems. This allowed them to study the flow of energy and material through ecological systems.

Processes

File:River gambia Niokolokoba National Park.gif|thumb|Rainforest ecosystems are rich in biodiversity. This is the Gambia River in Senegal's Niokolo-Koba National ParkNiokolo-Koba National ParkFile:Baja California Desert.jpg|thumb|Flora of Baja California Desert, Cataviña region, MexicoMexico(File:Vegetation.png|thumb|right|Biomes of the world)Ecosystems are controlled both by external and internal factors. External factors, also called state factors, control the overall structure of an ecosystem and the way things work within it, but are not themselves influenced by the ecosystem. The most important of these is climate.Chapin et al. (2002), pp. 11–13 Climate determines the biome in which the ecosystem is embedded. Rainfall patterns and seasonal temperatures influence photosynthesis and thereby determine the amount of water and energy available to the ecosystem.Parent material determines the nature of the soil in an ecosystem, and influences the supply of mineral nutrients. Topography also controls ecosystem processes by affecting things like microclimate, soil development and the movement of water through a system. For example, ecosystems can be quite different if situated in a small depression on the landscape, versus one present on an adjacent steep hillside.Other external factors that play an important role in ecosystem functioning include time and potential biota. Similarly, the set of organisms that can potentially be present in an area can also significantly affect ecosystems. Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present. The introduction of non-native species can cause substantial shifts in ecosystem function.JOURNAL, Simberloff, Daniel, Martin, Jean-Louis, Genovesi, Piero, Maris, Virginie, Wardle, David A., Aronson, James, Courchamp, Franck, Galil, Bella, García-Berthou, Emili, 2013, Impacts of biological invasions: what's what and the way forward, Trends in Ecology & Evolution, 28, 1, 58–66, 10.1016/j.tree.2012.07.013, 22889499, 0169-5347, Unlike external factors, internal factors in ecosystems not only control ecosystem processes but are also controlled by them. Consequently, they are often subject to feedback loops. While the resource inputs are generally controlled by external processes like climate and parent material, the availability of these resources within the ecosystem is controlled by internal factors like decomposition, root competition or shading. Other factors like disturbance, succession or the types of species present are also internal factors.{{cn|date=December 2018}}

Primary production

File:Seawifs global biosphere.jpg|upright=1.8|thumb|Global oceanic and terrestrial phototroph abundance, from September 1997 to August 2000. As an estimate of autotrophautotrophPrimary production is the production of organic matter from inorganic carbon sources. This mainly occurs through photosynthesis. The energy incorporated through this process supports life on earth, while the carbon makes up much of the organic matter in living and dead biomass, soil carbon and fossil fuels. It also drives the carbon cycle, which influences global climate via the greenhouse effect.Through the process of photosynthesis, plants capture energy from light and use it to combine carbon dioxide and water to produce carbohydrates and oxygen. The photosynthesis carried out by all the plants in an ecosystem is called the gross primary production (GPP).Chapin et al. (2002), pp. 97–104 About half of the GPP is consumed in plant respiration."Chapin et al. 2011, p. 157 The remainder, that portion of GPP that is not used up by respiration, is known as the net primary production (NPP).Chapin et al. (2002), pp. 123–150 Total photosynthesis is limited by a range of environmental factors. These include the amount of light available, the amount of leaf area a plant has to capture light (shading by other plants is a major limitation of photosynthesis), rate at which carbon dioxide can be supplied to the chloroplasts to support photosynthesis, the availability of water, and the availability of suitable temperatures for carrying out photosynthesis.

Energy flow

{{See also|Food web|Trophic level}}Energy and carbon enter ecosystems through photosynthesis, are incorporated into living tissue, transferred to other organisms that feed on the living and dead plant matter, and eventually released through respiration.The carbon and energy incorporated into plant tissues (net primary production) is either consumed by animals while the plant is alive, or it remains uneaten when the plant tissue dies and becomes detritus. In terrestrial ecosystems, roughly 90% of the net primary production ends up being broken down by decomposers. The remainder is either consumed by animals while still alive and enters the plant-based trophic system, or it is consumed after it has died, and enters the detritus-based trophic system.{{cn|date=December 2018}}In aquatic systems, the proportion of plant biomass that gets consumed by herbivores is much higher.Chapin et al. (2002) pp. 244–264In trophic systems photosynthetic organisms are the primary producers. The organisms that consume their tissues are called primary consumers or secondary producers—herbivores. Organisms which feed on microbes (bacteria and fungi) are termed microbivores. Animals that feed on primary consumers—carnivores—are secondary consumers. Each of these constitutes a trophic level.The sequence of consumption—from plant to herbivore, to carnivore—forms a food chain. Real systems are much more complex than this—organisms will generally feed on more than one form of food, and may feed at more than one trophic level. Carnivores may capture some prey which are part of a plant-based trophic system and others that are part of a detritus-based trophic system (a bird that feeds both on herbivorous grasshoppers and earthworms, which consume detritus). Real systems, with all these complexities, form food webs rather than food chains. The food chain usually consist of four level of consumption which are producers, primary consumers, secondary consumers, and tertiary consumers.

Decomposition

{{See also|Decomposition}}The carbon and nutrients in dead organic matter are broken down by a group of processes known as decomposition. This releases nutrients that can then be re-used for plant and microbial production and returns carbon dioxide to the atmosphere (or water) where it can be used for photosynthesis. In the absence of decomposition, the dead organic matter would accumulate in an ecosystem, and nutrients and atmospheric carbon dioxide would be depleted.Chapin et al. (2002), pp. 151–157 Approximately 90% of terrestrial net primary production goes directly from plant to decomposer.Decomposition processes can be separated into three categories—leaching, fragmentation and chemical alteration of dead material. As water moves through dead organic matter, it dissolves and carries with it the water-soluble components. These are then taken up by organisms in the soil, react with mineral soil, or are transported beyond the confines of the ecosystem (and are considered lost to it). Newly shed leaves and newly dead animals have high concentrations of water-soluble components and include sugars, amino acids and mineral nutrients. Leaching is more important in wet environments and much less important in dry ones.Fragmentation processes break organic material into smaller pieces, exposing new surfaces for colonization by microbes. Freshly shed leaf litter may be inaccessible due to an outer layer of cuticle or bark, and cell contents are protected by a cell wall. Newly dead animals may be covered by an exoskeleton. Fragmentation processes, which break through these protective layers, accelerate the rate of microbial decomposition. Animals fragment detritus as they hunt for food, as does passage through the gut. Freeze-thaw cycles and cycles of wetting and drying also fragment dead material.The chemical alteration of the dead organic matter is primarily achieved through bacterial and fungal action. Fungal hyphae produce enzymes which can break through the tough outer structures surrounding dead plant material. They also produce enzymes which break down lignin, which allows them access to both cell contents and to the nitrogen in the lignin. Fungi can transfer carbon and nitrogen through their hyphal networks and thus, unlike bacteria, are not dependent solely on locally available resources.Decomposition rates vary among ecosystems.JOURNAL, Ochoa-Hueso, R, Delgado-Baquerizo, M, King, PTA, Benham, M, Arca, V, Power, SA, Ecosystem type and resource quality are more important than global change drivers in regulating early stages of litter decomposition, Soil Biology and Biochemistry, February 2019, 129, 144–152, 10.1016/j.soilbio.2018.11.009, The rate of decomposition is governed by three sets of factors—the physical environment (temperature, moisture, and soil properties), the quantity and quality of the dead material available to decomposers, and the nature of the microbial community itself.Chapin et al. (2002), pp. 159–174 Temperature controls the rate of microbial respiration; the higher the temperature, the faster microbial decomposition occurs. It also affects soil moisture, which slows microbial growth and reduces leaching. Freeze-thaw cycles also affect decomposition—freezing temperatures kill soil microorganisms, which allows leaching to play a more important role in moving nutrients around. This can be especially important as the soil thaws in the spring, creating a pulse of nutrients which become available.Decomposition rates are low under very wet or very dry conditions. Decomposition rates are highest in wet, moist conditions with adequate levels of oxygen. Wet soils tend to become deficient in oxygen (this is especially true in wetlands), which slows microbial growth. In dry soils, decomposition slows as well, but bacteria continue to grow (albeit at a slower rate) even after soils become too dry to support plant growth.{{Further|Decomposition#Rate of decomposition}}

Nutrient cycling

{{See also|Nutrient cycle|Biogeochemical cycle|Nitrogen cycle}}(File:Nitrogen Cycle.jpg|thumb|right|upright=1.8|Biological nitrogen cycling)Ecosystems continually exchange energy and carbon with the wider environment. Mineral nutrients, on the other hand, are mostly cycled back and forth between plants, animals, microbes and the soil. Most nitrogen enters ecosystems through biological nitrogen fixation, is deposited through precipitation, dust, gases or is applied as fertilizer.Chapin et al. (2002), pp. 197–215Since most terrestrial ecosystems are nitrogen-limited, nitrogen cycling is an important control on ecosystem production.Until modern times, nitrogen fixation was the major source of nitrogen for ecosystems. Nitrogen-fixing bacteria either live symbiotically with plants or live freely in the soil. The energetic cost is high for plants which support nitrogen-fixing symbionts—as much as 25% of gross primary production when measured in controlled conditions. Many members of the legume plant family support nitrogen-fixing symbionts. Some cyanobacteria are also capable of nitrogen fixation. These are phototrophs, which carry out photosynthesis. Like other nitrogen-fixing bacteria, they can either be free-living or have symbiotic relationships with plants. Other sources of nitrogen include acid deposition produced through the combustion of fossil fuels, ammonia gas which evaporates from agricultural fields which have had fertilizers applied to them, and dust. Anthropogenic nitrogen inputs account for about 80% of all nitrogen fluxes in ecosystems.When plant tissues are shed or are eaten, the nitrogen in those tissues becomes available to animals and microbes. Microbial decomposition releases nitrogen compounds from dead organic matter in the soil, where plants, fungi, and bacteria compete for it. Some soil bacteria use organic nitrogen-containing compounds as a source of carbon, and release ammonium ions into the soil. This process is known as nitrogen mineralization. Others convert ammonium to nitrite and nitrate ions, a process known as nitrification. Nitric oxide and nitrous oxide are also produced during nitrification. Under nitrogen-rich and oxygen-poor conditions, nitrates and nitrites are converted to nitrogen gas, a process known as denitrification.Other important nutrients include phosphorus, sulfur, calcium, potassium, magnesium and manganese.Chapin et al. (2002), pp. 215–222 Phosphorus enters ecosystems through weathering. As ecosystems age this supply diminishes, making phosphorus-limitation more common in older landscapes (especially in the tropics). Calcium and sulfur are also produced by weathering, but acid deposition is an important source of sulfur in many ecosystems. Although magnesium and manganese are produced by weathering, exchanges between soil organic matter and living cells account for a significant portion of ecosystem fluxes. Potassium is primarily cycled between living cells and soil organic matter.

Function and biodiversity

{{See also|Ecosystem diversity}}File:View of loch lomond.JPG|thumb|Loch Lomond in Scotland forms a relatively isolated ecosystem. The fish community of this lake has remained stable over a long period until a number of introductions in the 1970s restructured it's (food web]].JOURNAL, Adams, C.E., The fish community of Loch Lomond, Scotland: its history and rapidly changing status, Hydrobiologia, 1994, 290, 1–3, 91–102, 10.1007/BF00008956, )File:Spiny Forest Ifaty Madagascar.jpg|thumb|Spiny forest at Ifaty, Madagascar, featuring various Adansonia (baobab) species, Alluaudia proceraAlluaudia proceraBiodiversity plays an important role in ecosystem functioning.Schulze et al. (2005), pp. 449–453 The reason for this is that ecosystem processes are driven by the number of species in an ecosystem, the exact nature of each individual species, and the relative abundance organisms within these species.Chapin et al. (2002), pp. 265–277 Ecosystem processes are broad generalizations that actually take place through the actions of individual organisms. The nature of the organisms—the species, functional groups and trophic levels to which they belong—dictates the sorts of actions these individuals are capable of carrying out and the relative efficiency with which they do so.{{cn|date=December 2018}}Ecological theory suggests that in order to coexist, species must have some level of limiting similarity—they must be different from one another in some fundamental way, otherwise one species would competitively exclude the other.BOOK, Schoener, Thomas W., The Princeton Guide to Ecology, Simon A. Levin, Princeton University Press, Princeton, 2009, 2–13, Ecological Niche, 978-0-691-12839-9, Despite this, the cumulative effect of additional species in an ecosystem is not linear—additional species may enhance nitrogen retention, for example, but beyond some level of species richness, additional species may have little additive effect.The addition (or loss) of species which are ecologically similar to those already present in an ecosystem tends to only have a small effect on ecosystem function. Ecologically distinct species, on the other hand, have a much larger effect. Similarly, dominant species have a large effect on ecosystem function, while rare species tend to have a small effect. Keystone species tend to have an effect on ecosystem function that is disproportionate to their abundance in an ecosystem. Similarly, an ecosystem engineer is any organism that creates, significantly modifies, maintains or destroys a habitat.{{cn|date=December 2018}}

Dynamics

Ecosystems are dynamic entities. They are subject to periodic disturbances and are in the process of recovering from some past disturbance.Chapin et al. (2002), pp. 281–304 When a perturbation occurs, an ecosystem responds by moving away from its initial state. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance, is termed its resistance. On the other hand, the speed with which it returns to its initial state after disturbance is called its resilience. Time plays a role in the development of soil from bare rock and the recovery of a community from disturbance.From one year to another, ecosystems experience variation in their biotic and abiotic environments. A drought, a colder than usual winter, and a pest outbreak all are short-term variability in environmental conditions. Animal populations vary from year to year, building up during resource-rich periods and crashing as they overshoot their food supply. These changes play out in changes in net primary production decomposition rates, and other ecosystem processes. Longer-term changes also shape ecosystem processes—the forests of eastern North America still show legacies of cultivation which ceased 200 years ago, while methane production in eastern Siberian lakes is controlled by organic matter which accumulated during the Pleistocene.Disturbance also plays an important role in ecological processes. F. Stuart Chapin and coauthors define disturbance as "a relatively discrete event in time and space that alters the structure of populations, communities, and ecosystems and causes changes in resources availability or the physical environment".Chapin et al. (2002), p. 285 This can range from tree falls and insect outbreaks to hurricanes and wildfires to volcanic eruptions. Such disturbances can cause large changes in plant, animal and microbe populations, as well soil organic matter content. Disturbance is followed by succession, a "directional change in ecosystem structure and functioning resulting from biotically driven changes in resources supply."The frequency and severity of disturbance determine the way it affects ecosystem function. A major disturbance like a volcanic eruption or glacial advance and retreat leave behind soils that lack plants, animals or organic matter. Ecosystems that experience such disturbances undergo primary succession. A less severe disturbance like forest fires, hurricanes or cultivation result in secondary succession and a faster recovery. More severe disturbance and more frequent disturbance result in longer recovery times.{{clear}}File:Panorama presa las niñas mogan gran canaria.jpg|thumb|upright=4.1|center|A freshwater lake in Gran Canaria, an island of the Canary IslandsCanary Islands

Ecosystem ecology

{{See also|Ecosystem model}}File:East Scotia Ridge - Plos Biol 04.tif|thumb|right|A hydrothermal venthydrothermal ventEcosystem ecology studies the processes and dynamics of ecosystems, and the way the flow of matter and energy through them structures natural systems. The study of ecosystems can cover 10 orders of magnitude, from the surface layers of rocks to the surface of the planet.Chapin et al. (2002), pp. 3–7There is no single definition of what constitutes an ecosystem.JOURNAL, Sagoff, Mark, 2003, The plaza and the pendulum: Two concepts of ecological science, Biology and Philosophy, 18, 4, 529–552, 10.1023/A:1025566804906, German ecologist Ernst-Detlef Schulze and coauthors defined an ecosystem as an area which is "uniform regarding the biological turnover, and contains all the fluxes above and below the ground area under consideration." They explicitly reject Gene Likens' use of entire river catchments as "too wide a demarcation" to be a single ecosystem, given the level of heterogeneity within such an area.Schulze et al. 300–402 Other authors have suggested that an ecosystem can encompass a much larger area, even the whole planet.Willis (1997), p. 269; Chapin et al. (2002), p. 5; Krebs (2009). p. 572 Schulze and coauthors also rejected the idea that a single rotting log could be studied as an ecosystem because the size of the flows between the log and its surroundings are too large, relative to the proportion cycles within the log. Philosopher of science Mark Sagoff considers the failure to define "the kind of object it studies" to be an obstacle to the development of theory in ecosystem ecology.Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation.JOURNAL, Carpenter, Stephen R., Jonathan J. Cole, Timothy E. Essington, James R. Hodgson, Jeffrey N. Houser, James F. Kitchell, Michael L. Pace, 1998, Evaluating Alternative Explanations in Ecosystem Experiments,weblink Ecosystems, 1, 4, 335–344, 10.1007/s100219900025, Studies can be carried out at a variety of scales, ranging from whole-ecosystem studies to studying (Microcosm: Model / experimental ecosystem|microcosms) or mesocosms (simplified representations of ecosystems).JOURNAL, Schindler, David W., 1998, Replication versus Realism: The Need for Ecosystem-Scale Experiments, Ecosystems, 1, 4, 323–334, 10.1007/s100219900026, 3658915, American ecologist Stephen R. Carpenter has argued that microcosm experiments can be "irrelevant and diversionary" if they are not carried out in conjunction with field studies done at the ecosystem scale. Microcosm experiments often fail to accurately predict ecosystem-level dynamics.JOURNAL, Carpenter, Stephen R., 1996, Microcosm Experiments have Limited Relevance for Community and Ecosystem Ecology, Ecology, 77, 3, 677–680, 10.2307/2265490, 2265490, The Hubbard Brook Ecosystem Study started in 1963 to study the White Mountains in New Hampshire. It was the first successful attempt to study an entire watershed as an ecosystem. The study used stream chemistry as a means of monitoring ecosystem properties, and developed a detailed biogeochemical model of the ecosystem.BOOK, Effective Ecological Monitoring, Lindenmayer, David B., Gene E. Likens, CSIRO Publishing, 2010, 978-1-84971-145-6, Collingwood, Australia, 87–145, The Problematic, the Effective and the Ugly – Some Case Studies, Long-term research at the site led to the discovery of acid rain in North America in 1972. Researchers documented the depletion of soil cations (especially calcium) over the next several decades.JOURNAL, Likens, Gene E., 2004, Some perspectives on long-term biogeochemical research from the Hubbard Brook Ecosystem Study,weblink yes, Ecology, 85, 9, 2355–2362, 10.1890/03-0243, 3450233,weblink" title="web.archive.org/web/20130501031912weblink">weblink 2013-05-01,

Human activities

Human activities are important in almost all ecosystems. Although humans exist and operate within ecosystems, their cumulative effects are large enough to influence external factors like climate.

Ecosystem goods and services

File:Avalanche Lake, looking south.jpg|thumb|The High Peaks Wilderness Area in the {{convert|6000000|acre|sing=on}} Adirondack ParkAdirondack Park{{See also|Ecosystem valuation|Ecological yield}}Ecosystems provide a variety of goods and services upon which people depend.JOURNAL, Christensen, Norman L., Ann M., Bartuska, James H., Brown, Stephen, Carpenter, Carla, D'Antonio, Robert, Francis, Jerry F., Franklin, James A., MacMahon, Reed F., Noss, David J., Parsons, Charles H., Peterson, Monica G., Turner, Robert G., Woodmansee, 1996, The Report of the Ecological Society of America Committee on the Scientific Basis for Ecosystem Management, Ecological Applications, 6, 3, 665–691, 10.2307/2269460, 10.1.1.404.4909, 2269460, Ecosystem goods include the "tangible, material products" of ecosystem processes such as food, construction material, medicinal plants.JOURNAL, Brown, Thomas C., John C. Bergstrom, John B. Loomis, 2007, Defining, valuing and providing ecosystem goods and services,weblink yes, Natural Resources Journal, 47, 2, 329–376,weblink" title="web.archive.org/web/20130525213257weblink">weblink 2013-05-25, They also include less tangible items like tourism and recreation, and genes from wild plants and animals that can be used to improve domestic species.Ecosystem services, on the other hand, are generally "improvements in the condition or location of things of value". These include things like the maintenance of hydrological cycles, cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. While material from the ecosystem had traditionally been recognized as being the basis for things of economic value, ecosystem services tend to be taken for granted.

Ecosystem management

{{See also|Ecological economics|Sustainability|Sustainable development}}When natural resource management is applied to whole ecosystems, rather than single species, it is termed ecosystem management.JOURNAL, Grumbine, R. Edward, 1994, What is ecosystem management?, Conservation Biology, 8, 1, 27–38,weblink 10.1046/j.1523-1739.1994.08010027.x, yes,weblink" title="web.archive.org/web/20130502051519weblink">weblink 2013-05-02, Although definitions of ecosystem management abound, there is a common set of principles which underlie these definitions.BOOK, Principles of Terrestrial Ecosystem Ecology, Chapin, F. Stuart, Pamela A. Matson, Harold A. Mooney, Springer, 2002, 978-0-387-95443-1, New York, 362–365, A fundamental principle is the long-term sustainability of the production of goods and services by the ecosystem; "intergenerational sustainability [is] a precondition for management, not an afterthought".While ecosystem management can be used as part of a plan for wilderness conservation, it can also be used in intensively managed ecosystems (see, for example, agroecosystem and close to nature forestry).

Threats caused by humans

{{see also|Planetary boundaries}}As human population and per capita consumption grow, so do the resource demands imposed on ecosystems and the effects of the human ecological footprint. Natural resources are vulnerable and limited. The environmental impacts of anthropogenic actions are becoming more apparent. Problems for all ecosystems include: environmental pollution, climate change and biodiversity loss. For terrestrial ecosystems further threats include air pollution, soil degradation, and deforestation. For aquatic ecosystems threats include also unsustainable exploitation of marine resources (for example overfishing of certain species), marine pollution, microplastics pollution, water pollution, the warming of oceans, and building on coastal areas.BOOK, Alexander, David E., Encyclopedia of Environmental Science, Springer Science+Business Media, Springer, 1 May 1999, 978-0-412-74050-3, Society is increasingly becoming aware that ecosystem services are not only limited but also that they are threatened by human activities. The need to better consider long-term ecosystem health and its role in enabling human habitation and economic activity is urgent. To help inform decision-makers, many ecosystem services are being assigned economic values, often based on the cost of replacement with anthropogenic alternatives. The ongoing challenge of prescribing economic value to nature, for example through biodiversity banking, is prompting transdisciplinary shifts in how we recognize and manage the environment, social responsibility, business opportunities, and our future as a species.{{citation needed|date=March 2018}}

See also

Notes

{{reflist|group=fn}}

References

{{Reflist}}

Literature cited

  • BOOK, Chapin, F. Stuart, Pamela A. Matson, Harold A. Mooney, Principles of Terrestrial Ecosystem Ecology, Springer, New York, 2002, 978-0-387-95443-1,
  • BOOK, Chapin, F. Stuart, Pamela A. Matson, Peter M. Vitousek, Principles of Terrestrial Ecosystem Ecology, Second, Springer, New York, 2011, 978-1-4419-9503-2,
  • BOOK, Gurevitch, Jessica, Samuel M. Scheiner, Gordon A. Fox, The Ecology of Plants, Sinauer Associates, Sunderland, Massachusetts, 2006, Second, 978-0-87893-294-8,
  • BOOK, Krebs, Charles J., Ecology: The Experimental Analysis of Distribution and Abundance, Benjamin Cummings, San Francisco, 2009, Sixth, 978-0-321-50743-3,
  • BOOK, Molles, Manuel C., Ecology: Concepts and Applications, WCB/McGraw-HIll, Boston, 1999, 978-0-07-042716-7,
  • BOOK, Schulze, Ernst-Detlef, Erwin Beck, Klaus Müller-Hohenstein, Plant Ecology, Springer, Berlin, 2005, 978-3-540-20833-4,
  • BOOK, Smith, Thomas M., Robert Leo Smith, Elements of Ecology, Benjamin Cummings, Boston, 2012, Eighth, 978-0-321-73607-9,
  • JOURNAL, 10.2307/1930070, Tansley, AG, 1935, The use and abuse of vegetational terms and concepts, Ecology, 16, 3, 284–307, 1930070, weblink" title="web.archive.org/web/20161006125220weblink">PDF.
  • JOURNAL, Willis, A.J., 1997, The Ecosystem: An Evolving Concept Viewed Historically, Functional Ecology, 11, 2, 268–271, 10.1111/j.1365-2435.1997.00081.x,

External links

{{Commons category|Ecosystems}}{{Wiktionary}}{{Scholia|topic}} {{modelling ecosystems}}{{Composition (Biology)}}{{Physical Earth}}{{Nature nav}}{{Systems}}{{Authority control}}

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