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bear
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{{about|the carnivoran mammals}}{{pp-move-indef}}{{pp-semi|small=yes||expiry=}}{{short description|Family of mammals}}{{good article}}{{Automatic taxobox| taxon=Ursidae| name=Bears
38|0}}Late Eocene – Recent| image=2010-kodiak-bear-1.jpg| image_caption=Brown bear in AlaskaJohann Fischer von Waldheim>G. Fischer de Waldheim, 1817| subdivision_ranks=Subfamilies| subdivision ={{dagger}}Amphicynodontinae{{dagger}}Hemicyoninae{{dagger}}Ursavinae{{dagger}}AgriotheriinaeAiluropodinaeTremarctinaeUrsinae}}Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Northern Hemisphere and partially in the Southern Hemisphere. Bears are found on the continents of North America, South America, Europe, and Asia. Common characteristics of modern bears include large bodies with stocky legs, long snouts, small rounded ears, shaggy hair, plantigrade paws with five nonretractile claws, and short tails.While the polar bear is mostly carnivorous, and the giant panda feeds almost entirely on bamboo, the remaining six species are omnivorous with varied diets. With the exception of courting individuals and mothers with their young, bears are typically solitary animals. They may be diurnal or nocturnal and have an excellent sense of smell. Despite their heavy build and awkward gait, they are adept runners, climbers, and swimmers. Bears use shelters, such as caves and logs, as their dens; most species occupy their dens during the winter for a long period of hibernation, up to 100 days.Bears have been hunted since prehistoric times for their meat and fur; they have been used for bear-baiting and other forms of entertainment, such as being made to dance. With their powerful physical presence, they play a prominent role in the arts, mythology, and other cultural aspects of various human societies. In modern times, bears have come under pressure through encroachment on their habitats and illegal trade in bear parts, including the Asian bile bear market. The IUCN lists six bear species as vulnerable or endangered, and even least concern species, such as the brown bear, are at risk of extirpation in certain countries. The poaching and international trade of these most threatened populations are prohibited, but still ongoing.

Etymology

The English word "bear" comes from Old English ' and belongs to a family of names for the bear in Germanic languages, such as Swedish ', also used as a first name. This form is conventionally said to be related to a Proto-Indo-European word for "brown", so that "bear" would mean "the brown one".BOOK,weblink The Oxford Iintroduction to Proto Indo European and the Proto Indo European World, Mallory, J. P., Adams, Douglas Q., 2006, Oxford University Press, 9781429471046, New York, 333, 139999117, BOOK, Indo-European Language and Culture: An Introduction., Fortson, Benjamin W., 2011, John Wiley & Sons, 9781444359688, 2nd, Hoboken, 31, 778339290, However, Ringe notes that while this etymology is semantically plausible, a word meaning "brown" of this form cannot be found in Proto-Indo-European. He suggests instead that "bear" is from the Proto-Indo-European word *ǵʰwḗr- ~ *ǵʰwér "wild animal".BOOK,weblink From Proto-Indo-European to Proto-Germanic, Ringe, Don, Oxford University Press, 2017, 9780192511188, 2nd, A Linguistic History of English, 1, Oxford, 128, This terminology for the animal originated as a taboo avoidance term: proto-Germanic tribes replaced their original word for bear—arkto—with this euphemistic expression out of fear that speaking the animal's true name might cause it to appear.DICTIONARY, bear (n.),weblink Online Etymology Dictionary, 22 January 2017,weblink" title="web.archive.org/web/20170202020051weblink">weblink 2017-02-02, live, WEB, Postma, Laura, The word for "bear",weblink University of Pittsburgh Slovak Studies Program, 21 March 2018,weblink" title="web.archive.org/web/20171122151051weblink">weblink 2017-11-22, live, According to author Ralph Keyes, this is the oldest known euphemism.NEWS, Silver, Alexandra, Hooking Up and Using the John: Why Do We Use So Many Euphemisms?,weblink 4 April 2019, Time, 10 January 2011, Bear taxon names such as Arctoidea and Helarctos come from the ancient Greek word ἄρκτος (arktos), meaning bear,DICTIONARY, Liddell, Henry George, Scott, Robert,weblink Arktos, A Greek-English Lexicon, Perseus Digital Library, as do the names "arctic" and "antarctic", from the constellation Ursa Major, the "Great Bear", prominent in the northern sky.WEB,weblink The Great Bear Constellation Ursa Major, 12 January 2017,weblink" title="web.archive.org/web/20101130031637weblink">weblink 30 November 2010, live, Bear taxon names such as Ursidae and Ursus come from Latin Ursus/Ursa, he-bear/she-bear. The female first name "Ursula", originally derived from a Christian saint's name, means "little she-bear" (diminutive of Latin ursa). In Switzerland, the male first name "Urs" is especially popular, while the name of the canton and city of Bern is derived from Bär, German for bear. The Germanic name Bernard (including Bernhardt and similar forms) means "bear-brave", "bear-hardy", or "bold bear".WEB,weblink Ursa Major – the Greater Bear, constellationsofwords.com, 11 March 2015,weblink" title="web.archive.org/web/20150314234800weblink">weblink 2015-03-14, live, WEB,weblink Bernhard Family History, ancestry.com, 11 March 2015,weblink" title="web.archive.org/web/20150402201212weblink">weblink 2015-04-02, live, The Old English name Beowulf is a kenning, "bee-wolf", for bear, in turn meaning a brave warrior.Sweet, Henry (1884) Anglo-Saxon Reader in Prose and Verse. The Clarendon Press, p. 202.

Taxonomy

{{see also|List of bears}}The family Ursidae is one of nine families in the suborder Caniformia, or "doglike" carnivorans, within the order Carnivora. Bears' closest living relatives are the pinnipeds, canids, and musteloids.JOURNAL, Welsey-Hunt, G. D., Flynn, J. J., 2005, Phylogeny of the Carnivora: basal relationships among the Carnivoramorphans, and assessment of the position of 'Miacoidea' relative to Carnivora, Journal of Systematic Palaeontology, 3, 1, 1–28, 10.1017/S1477201904001518, Modern bears comprise eight species in three subfamilies: Ailuropodinae (monotypic with the giant panda), Tremarctinae (monotypic with the spectacled bear), and Ursinae (containing six species divided into one to three genera, depending on the authority). Nuclear chromosome analysis show that the karyotype of the six ursine bears is nearly identical, with each having 74 chromosomes, whereas the giant panda has 42 chromosomes and the spectacled bear 52. These smaller numbers can be explained by the fusing of some chromosomes, and the banding patterns on these match those of the ursine species, but differ from those of procyonids, which supports the inclusion of these two species in Ursidae rather than in Procyonidae, where they had been placed by some earlier authorities.BOOK, Servheen, C., Herrero, S., Peyton, B., Bears: Status Survey and Conservation Action Plan,weblink 1999, IUCN, 978-2-8317-0462-3, 26–30,

Evolution

File:Plithocyon armagnacensis.JPG|thumb|left|Plithocyon armagnacensis skull, a member of the extinct subfamily Hemicyoninae from the MioceneMioceneThe earliest members of Ursidae belong to the extinct subfamily Amphicynodontinae, including Parictis (late Eocene to early middle Miocene, 38–18 Mya) and the slightly younger Allocyon (early Oligocene, 34–30 Mya), both from North America. These animals looked very different from today's bears, being small and raccoon-like in overall appearance, with diets perhaps more similar to that of a badger. Parictis does not appear in Eurasia and Africa until the Miocene.BOOK, Kemp, T. S., 2005, The Origin and Evolution of Mammals, Oxford University Press, 978-0-19-850760-4, 260, It is unclear whether late-Eocene ursids were also present in Eurasia, although faunal exchange across the Bering land bridge may have been possible during a major sea level low stand as early as the late Eocene (about 37 Mya) and continuing into the early Oligocene.JOURNAL, Banyue, Wang, Zhanxiang, Qiu, 2005, Notes on Early Oligocene Ursids (Carnivora, Mammalia) from Saint Jacques, Nei Mongol, China, Bulletin of the American Museum of Natural History, 279, 116–124,weblink 10.1206/0003-0090(2003)2792.0.CO;2, 279,weblink" title="web.archive.org/web/20090326201348weblink">weblink 2009-03-26, live, European genera morphologically very similar to Allocyon, and to the much younger American Kolponomos (about 18 Mya),JOURNAL, Tedford, R. H., Richard H. Tedford, Barnes, L. G., Ray, C. E., The early Miocene littoral ursoid carnivoran Kolponomos: Systematics and mode of life, 1994, Proceedings of the San Diego Society of Natural History, 29, 11–32,weblinkweblink" title="web.archive.org/web/20120922142556weblink">weblink dead, 2012-09-22, are known from the Oligocene, including Amphicticeps and Amphicynodon. There has been various morphological evidence linking amphicynodontines with pinnipeds, as both groups were semi-aquatic, otter-like mammals.JOURNAL, Tedford, R. H., Richard H. Tedford, Barnes, L. G., Ray, C. E., The early Miocene littoral ursoid carnivoran Kolponomos: Systematics and mode of life, 1994, Proceedings of the San Diego Society of Natural History, 29, 11–32,weblinkweblink" title="web.archive.org/web/20120922142556weblink">weblink dead, 22 September 2012, 24 July 2010, JOURNAL, Rybczynski, N., Dawson, M.R., Tedford, R.H., 2009, A semi-aquatic Arctic mammalian carnivore from the Miocene epoch and origin of Pinnipedia, Nature, 458, 7241, 1021–24, 10.1038/nature07985, 19396145, 2009Natur.458.1021R, JOURNAL, Berta, A., Morgan, C., Boessenecker, R.W., 2018, The Origin and Evolutionary Biology of Pinnipeds: Seals, Sea Lions, and Walruses, Annual Review of Earth and Planetary Sciences, 0, 203–228, 10.1146/annurev-earth-082517-010009, 2018AREPS..46..203B, In addition to the support of the pinniped–amphicynodontine clade, other morphological and some molecular evidence supports bears being the closet living relatives to pinnipeds.JOURNAL, Hunt, R. M. Jr., Barnes, L. G., 1994, Basicranial evidence for ursid affinity of the oldest pinnipeds, Proceedings of the San Diego Society of Natural History, 29, 57–67,weblink PDF, JOURNAL, Lento, G. M., Hickson, R. E., Chambers, G. K., Penny, D., 1995,weblink Use of spectral analysis to test hypotheses on the origin of pinnipeds, Molecular Biology and Evolution, 12, 1, 28–52, 7877495, 10.1093/oxfordjournals.molbev.a040189, 2018-04-03,weblink" title="web.archive.org/web/20081007185649weblink">weblink 2008-10-07, live, JOURNAL, Wang, X., McKenna, M. C., Dashzeveg, D., Amphicticeps and Amphicynodon (Arctoidea, Carnivora) from Hsanda Gol Formation, central Mongolia and phylogeny of basal arctoids with comments on zoogeography., 10.1206/0003-0082(2005)483[0001:AAAACF]2.0.CO;2, American Museum Novitates, 483, 3483, 216, 2005, 2246/5674, JOURNAL, Higdon, J. W., Bininda-Emonds, O. R., Beck, R. M., Ferguson, S. H., Phylogeny and divergence of the pinnipeds (Carnivora: Mammalia) assessed using a multigene dataset, 10.1186/1471-2148-7-216, BMC Evolutionary Biology, 7, 216, 2007, 17996107, 2245807, File:Raiola-manda-y-no-el-panda.jpg|thumb|right|Giant panda resting. This species is the most basal of the extant bears.]]The raccoon-sized, dog-like Cephalogale is the oldest-known member of the subfamily Hemicyoninae, which first appeared during the middle Oligocene in Eurasia about 30 Mya. The subfamily includes the younger genera Phoberocyon (20–15 Mya), and Plithocyon (15–7 Mya). A Cephalogale-like species gave rise to the genus Ursavus during the early Oligocene (30–28 Mya); this genus proliferated into many species in Asia and is ancestral to all living bears. Species of Ursavus subsequently entered North America, together with Amphicynodon and Cephalogale, during the early Miocene (21–18 Mya). Members of the living lineages of bears diverged from Ursavus between 15 and 20 Mya,JOURNAL, Waits, Lisette, Rapid radiation events in the family Ursidae indicated by likelihood phylogenetic estimation from multiple fragments of mtDNA, Molecular Phylogenetics and Evolution, 1999, 13, 1, 82–92,weblinkweblink" title="web.archive.org/web/20150903211855weblink">weblink dead, 2015-09-03, 10.1006/mpev.1999.0637, 10508542, JOURNAL, Pàges, Marie, Combined analysis of fourteen nuclear genes refines the Ursidae phylogeny, Molecular Phylogenetics and Evolution, 2008, 47, 1, 73–83, 10.1016/j.ympev.2007.10.019, 18328735, likely via the species Ursavus elmensis. Based on genetic and morphological data, the Ailuropodinae (pandas) were the first to diverge from other living bears about 19 Mya, although no fossils of this group have been found before about 5 Mya.JOURNAL, Krause, J., Unger, T., Noçon, A., Malaspinas, A., Kolokotronis, S., Stiller, M., Soibelzon, L., Spriggs, H., Dear, P. H., Briggs, A. W., Bray, S. C. E., O'Brien, S. J., Rabeder, G., Matheus, P., Cooper, A., Slatkin, M., Pääbo, S., Hofreiter, M., Mitochondrial genomes reveal an explosive radiation of extinct and extant bears near the Miocene-Pliocene boundary, BMC Evolutionary Biology, 8, 220, 220, 2008, 18662376, 2518930, 10.1186/1471-2148-8-220, The New World short-faced bears (Tremarctinae) differentiated from Ursinae following a dispersal event into North America during the mid-Miocene (about 13 Mya). They invaded South America (≈2.5 or 1.2 Ma) following formation of the Isthmus of Panama.JOURNAL, Soibelzon, L. H., Tonni, E. P., Bond, M., The fossil record of South American short-faced bears (Ursidae, Tremarctinae), Journal of South American Earth Sciences, 20, 1–2, 105–113, 2005, 10.1016/j.jsames.2005.07.005, 10915/5366, 2005JSAES..20..105S, Their earliest fossil representative is Plionarctos in North America (~ 10–2 Ma). This genus is probably the direct ancestor to the North American short-faced bears (genus Arctodus), the South American short-faced bears (Arctotherium), and the spectacled bears, Tremarctos, represented by both an extinct North American species (T. floridanus), and the lone surviving representative of the Tremarctinae, the South American spectacled bear (T. ornatus).File:Teufelshöhle-Höhlenbär-Dreiviertelprofil.jpg|thumb|Fossil of the cave bear (Ursus spelaeus), a relative of the brown bear and polar bear from the Pleistocene epoch in Europe]]The subfamily Ursinae experienced a dramatic proliferation of taxa about 5.3–4.5 Mya, coincident with major environmental changes; the first members of the genus Ursus appeared around this time. The sloth bear is a modern survivor of one of the earliest lineages to diverge during this radiation event (5.3 Mya); it took on its peculiar morphology, related to its diet of termites and ants, no later than by the early Pleistocene. By 3–4 Mya, the species Ursus minimus appears in the fossil record of Europe; apart from its size, it was nearly identical to today's Asian black bear. It is likely ancestral to all bears within Ursinae, perhaps aside from the sloth bear. Two lineages evolved from U. minimus: the black bears (including the sun bear, the Asian black bear, and the American black bear); and the brown bears (which includes the polar bear). Modern brown bears evolved from U. minimus via Ursus etruscus, which itself is ancestral to the extinct Pleistocene cave bear. Species of Ursinae have migrated repeatedly into North America from Eurasia as early as 4 Mya during the early Pliocene.JOURNAL, Qiu Zhanxiang, 2003, Dispersals of Neogene Carnivorans between Asia and North America, Bulletin of the American Museum of Natural History, 279, 18–31,weblink 10.1206/0003-0090(2003)2792.0.CO;2, 279,weblink" title="web.archive.org/web/20090326201351weblink">weblink 26 March 2009, live, Ward and Kynaston, pp. 74–77 The polar bear is the most recently evolved species and descended a population of brown bears that became isolated in northern latitudes by glaciation 400,000 years ago.JOURNAL, Liu, Shiping, Lorenzen, Eline D., Fumagalli, Matteo, Li, Bo, Harris, Kelley, Xiong, Zijun, Zhou, Long, Korneliussen, Thorfinn Sand, Somel, Mehmet, Babbitt, Courtney, Wray, Greg, Li, Jianwen, He, Weiming, Wang, Zhuo, Fu, Wenjing, Xiang, Xueyan, Morgan, Claire C., Doherty, Aoife, O'Connell, Mary J., McInerney, James O., Born, Erik W., Dalén, Love, Dietz, Rune, Orlando, Ludovic, Sonne, Christian, Zhang, Guojie, Nielsen, Rasmus, Willerslev, Eske, Wang, Jun, Population Genomics Reveal Recent Speciation and Rapid Evolutionary Adaptation in Polar Bears, Cell, 2014, 157, 4, 785–794, 10.1016/j.cell.2014.03.054,weblink 24813606, 4089990,

Phylogeny

The bears form a clade within the Carnivora. The cladogram is based on molecular phylogeny of six genes in Flynn, 2005.JOURNAL, Flynn, J. J., Finarelli, J. A., Zehr, S., Hsu, J., Nedbal, M. A., Molecular phylogeny of the Carnivora (Mammalia): Assessing the impact of increased sampling on resolving enigmatic relationships, 10.1080/10635150590923326, Systematic Biology, 54, 2, 317–337, 2005, 16012099, {{clade|label1=Carnivora|1={{clade
|1=Feliformia (File:Lydekker - Ocelot (white background).JPG|60px|Ocelot)
|label2=   Caniformia   
|2={{clade
|1=Canidae (File:Dogs, jackals, wolves, and foxes (Plate XI).jpg|60px|African golden wolf)
|label2=   Arctoidea   
|2={{clade
|label1=   
|1={{clade
|1=†Hemicyonidae (File:Hemicyon white background.jpg|60px|Hemicyon sansaniensis)
|2=Ursidae (File:Ursus arctos - 1700-1880 - Print - Iconographia Zoologica - Special Collections University of Amsterdam - (white background).jpg|70px|Brown bear)
}}
|2={{clade
|1=Pinnipedia (File:Faroe stamp 227 grey seal (Phoca vitulina) white background.jpg|60px|Common seal)
|label2=Musteloidea
|2={{clade
|1=Ailuridae (File:Ailurus fulgens - 1700-1880 - Print - Iconographia Zoologica - Special Collections University of Amsterdam -(white background).jpg|60px|Red panda)
|2=Procyonidae (File:Wild animals of North America, intimate studies of big and little creatures of the mammal kingdom (Page 410) (white background).jpg|60 px|Common raccoon)
|3=Mustelidae (File:Mustela eversmanii (white background).png|60px|Steppe polecat)
}}
}}
}}
}}
}}
}}Note that although they are called "bears" in some languages, red pandas and raccoons and their close relatives are not bears, but rather musteloids.There are two phylogenetic hypotheses on the relationships among extant and fossil bear species. One is all species of bears are classified in seven subfamilies as adopted here and related articles: Amphicynodontinae, Hemicyoninae, Ursavinae, Agriotheriinae, Ailuropodinae, Tremarctinae, and Ursinae.JOURNAL, McLellan, B., Reiner, D.C., 1992, A review of bear evolution, International Association for Bear Research and Management, 9, 1, 85–96, 10.2307/3872687, 3872687, JOURNAL, L. de Bonis, 2011, A new species of Adelpharctos (Mammalia, Carnivora, Ursidae) from the late Oligocene of the "Phosphorites du Quercy" (France), 10.3989/egeol.40553.181, Estudios Geológicos, 67, 2, 179–186, JOURNAL, Louis De Bonis, 2013, Ursidae (Mammalia, Carnivora) from the Late Oligocene of the "Phosphorites du Quercy" (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862, Geodiversitas, 35, 4, 787–814, 10.5252/g2013n4a4, JOURNAL, Qiu, Zhan-Xiang, etal, A Late Miocene Ursavus skull from Guanghe, Gansu, China, Vertebrata PalAsiatica, 52, 3, 2014, 265–302, Below is a cladogram of the subfamilies of bears after McLellan and Reiner (1992) and Qiu et a. (2014):{hide}clade| style=font-size:105%; line-height:105%;|label1=Ursidae|1={{Clade
|1={{dagger{edih}Amphicynodontinae (File:Kolponomos newportensis .jpg|75px)
|2={{Clade
|1={{dagger}}Hemicyoninae (File:Hemicyon white background.jpg|75px|Hemicyon sansaniensis)
|2={{Clade
|1={{dagger}}Ursavinae
|2={{Clade
|1={{dagger}}Agriotheriinae
|2={{Clade
|1=Ailuropodinae (File:Recherches pour servir à l'histoire naturelle des mammifères (Pl. 50) (white background).jpg|75px)
|2={{Clade
|1=Tremarctinae (File:Spectacled bear (1829).jpg|75px)
|2=Ursinae (File:Ursus arctos - 1700-1880 - Print - Iconographia Zoologica - Special Collections University of Amsterdam - (white background).jpg|75px)}} }} }} }} }} }} }}
The second alternative phylogenetic hypothesis was implemented by McKenna et al. (1997) is to classify all the bear species into the superfamily Ursoidea, with Hemicyoninae and Agriotheriinae being classified in the family "Hemicyonidae".BOOK, McKenna, M.C., Bell, S., 1997, Classification of Mammals Above the Species Level., Columbia University Press, New York, Amphicynodontinae under this classification were classified as stem-pinnipeds in the superfamily Phocoidea. In the McKenna and Bell classification both bears and pinnipeds in a parvorder of carnivoran mammals known as Ursida, along with the extinct bear dogs of the family Amphicyonidae. Below is the cladogram based on McKenna and Bell (1997) classification:{hide}clade| style=font-size:100%; line-height:100%;|label1=Ursida|1={{Clade
|1={{dagger{edih}Amphicyonidae (File:Daphoenodon superbus by R. B. Horsfall (coloured).png|75px)
|2={{Clade
|label1=Phocoidea
|1={{Clade
|1={{dagger}}Amphicynodontidae (File:Kolponomos newportensis .jpg|75px)
|2=Pinnipedia (File:Faroe stamp 227 grey seal (Phoca vitulina) white background.jpg|70px|Common seal)
}}
|label2=Ursoidea
|2={{Clade
|label1={{nowrap|{{dagger}}Hemicyoninae}}
|1={{Clade
|1={{dagger}}Hemicyoninae (File:Hemicyon white background.jpg|75px|Hemicyon sansaniensis)
|2={{dagger}}Agriotheriinae
}}
|label2=Ursidae
|2={{Clade
|1={{dagger}}Ursavinae
|2={{Clade
|1=Ailuropodinae (File:Recherches pour servir à l'histoire naturelle des mammifères (Pl. 50) (white background).jpg|75px)
|2={{Clade
|label1=
|1={{Clade
|1=Tremarctinae (File:Spectacled bear (1829).jpg|75px)
|2=Ursinae (File:Ursus arctos - 1700-1880 - Print - Iconographia Zoologica - Special Collections University of Amsterdam - (white background).jpg|75px)
}}
}}
}}
}}
}}
}}
}}
}}The phylogeny of extant bear species is shown in a cladogram based on complete mitochondrial DNA sequences from Yu et al., 2007.JOURNAL, Yu, Li, Li, Yi-Wei, Ryder, Oliver A., Zhang, Ya-Ping, Analysis of complete mitochondrial genome sequences increases phylogenetic resolution of bears (Ursidae), a mammalian family that experienced rapid speciation, BMC Evolutionary Biology, 2007, 7, 198, 198, 10.1186/1471-2148-7-198, 17956639, 2151078, The giant panda, followed by the spectacled bear are clearly the oldest species. The relationships of the other species are not very well resolved, though the polar bear and the brown bear form a close grouping.{{clade
|label1=Ursidae
|1={{clade
|1={{clade
|1={{clade
|1={{clade
|1={{clade
|1=Brown bear (File:Ursus arctos - 1700-1880 - Print - Iconographia Zoologica - Special Collections University of Amsterdam - (white background).jpg|70px)
|2=Polar bear (File:Lossy-page1-2518px-Ursus maritimus - 1700-1880 - Print - Iconographia Zoologica - Special Collections University of Amsterdam - (white background).jpg|70px)
}}
|2={{clade
|1={{clade
|1=Asian black bear (File:Ursus thibetanus - 1700-1880 - Print - Iconographia Zoologica - Special Collections University of Amsterdam -(white background).jpg|70px)
|2=American black bear (File:Ursus americanus - 1700-1880 - Print - Iconographia Zoologica - Special Collections University of Amsterdam - (white background).jpg|70px)
}}
|2=Sun bear (File:Ursus malayanus - 1700-1880 - Print - Iconographia Zoologica - Special Collections University of Amsterdam - (white background).jpg|70px)
}}
}}
|2=Sloth bear (File:Tremarctos ornatus 1824 (flipped).jpg|70px)
}}
|2=Spectacled bear (File:Spectacled bear (1829).jpg|70px)
}}
|2=Giant panda (File:Recherches pour servir à l'histoire naturelle des mammifères (Pl. 50) (white background).jpg|70px)
}}
}}

Physical characteristics

Size

{{multiple image
| direction = horizontal
| width = 200
| align = right
| image1 = Polar Bear AdF.jpg
| image2 = Sepilok Sabah BSBCC-photos-by-Wong-Siew-Te-06.jpg
| footer = Polar bear (left) and sun bear, the largest and smallest species respectively, on average
}}The bear family includes the most massive extant terrestrial members of the order Carnivora.{{efn|Treating pinnipedsBOOK, Illiger, J. K. W., Prodromus Systematis Mammalium et Avium, 1811, Sumptibus C. Salfeld, 138–39,weblink Latin, as marine mammals}} The polar bear is considered to be the largest extant species,Ward and Kynaston, p. 61 with adult males weighing {{convert|350|-|700|kg|0|abbr=on}} and measuring {{convert|2.4|-|3|m|ftin|sigfig=1}} in total length.BOOK, Hemstock, The Polar Bear, Hemstock, Annie, 1999, Capstone Press, Manakato, MN, 978-0-7368-0031-0, 4,weblink The Kodiak bear – the largest subspecies of the brown bear – roughly matches the polar bear in size and weight,Mark Carwardine,: Animal Records.. Sterling Publishing, 2008, {{ISBN|9781402756238}}, pp. 32 while another source states that the male Kodiak bear is the largest bear in the world, measuring up to {{convert|10|ft|m}} in standing height and {{convert|1500|lb|kg}} in weight.WEB,weblink Kodiak Bear Fact Sheet, Alaska Department of Fish and Game, Larry Van Daele, 2019, 13 September 2019, The smallest species is the sun bear, which ranges {{convert|25|–|65|kg|abbr=on}} in weight and {{convert|100|–|140|cm|abbr=on}} in length.JOURNAL, Fitzgerald, C. S., Krausman, P. S., 2002, Helarctos malayanus, Mammalian Species, 696, 1–5, 10.1644/1545-1410(2002)6962.0.CO;2, Prehistoric North and South American short-faced bears were the largest species known to have lived. The latter estimated to have weighed {{convert|1,600|kg|abbr=on}} and stood {{convert|3.4|m|abbr=on}} tall.{{citation |last=Dell'Amore |first=C. |year=2011 |url=http://news.nationalgeographic.com/news/2011/02/110203-biggest-bear-largest-giant-short-faced-animals-science/ |title=Biggest Bear Ever Found |archive-url=https://web.archive.org/web/20171017130448weblink |archive-date=2017-10-17 |publisher=National Geographic News |date=February 3, 2011}}JOURNAL, Soibelzon, L. H., Schubert, B. W., The Largest Known Bear, Arctotherium angustidens, from the Early Pleistocene Pampean Region of Argentina: With a Discussion of Size and Diet Trends in Bears, Journal of Paleontology, 85, 1, 69–75, January 2011,weblink 10.1666/10-037.1, 2011-06-01,weblink" title="web.archive.org/web/20110310053427weblink">weblink 2011-03-10, live, Body weight varies throughout the year in bears of temperate and arctic climates, as they build up fat reserves in the summer and autumn and lose weight during the winter.

Morphology

File:Bear foot.jpg|thumb|left|upright=1.2|Unlike most other Carnivora, bears have plantigrade feet. Drawing by Richard OwenRichard OwenBears are generally bulky and robust animals with short tails. They are sexually dimorphic with regard to size, with males typically being larger.JOURNAL, Derocher, Andrew E., Andersen, Magnus, Wiig, Øystein, Sexual dimorphism of polar bears, Journal of Mammalogy, 2005, 86, 5, 895–901, 10.1644/1545-1542(2005)86[895:SDOPB]2.0.CO;2,weblink dead,weblink" title="web.archive.org/web/20140202094143weblink">weblink 2014-02-02, BOOK, Hunt, R. M. Jr., 1998, Ursidae, Janis, Christine M., Scott, Kathleen M., Jacobs, Louis L., Evolution of Tertiary Mammals of North America, volume 1: Terrestrial carnivores, ungulates, and ungulatelike mammals, Cambridge, England, Cambridge University Press, 174–195, 978-0-521-35519-3, Larger species tend to show increased levels of sexual dimorphism in comparison to smaller species. Relying as they do on strength rather than speed, bears have relatively short limbs with thick bones to support their bulk. The shoulder blades and the pelvis are correspondingly massive. The limbs are much straighter than those of the big cats as there is no need for them to flex in the same way due to the differences in their gait. The strong forelimbs are used to catch prey, to excavate dens, to dig out burrowing animals, to turn over rocks and logs to locate prey, and to club large creatures.File:Black bear large.jpg|thumb|upright|Despite being quadrupedquadrupedUnlike most other land carnivorans, bears are plantigrade. They distribute their weight toward the hind feet, which makes them look lumbering when they walk. They are capable of bursts of speed but soon tire, and as a result mostly rely on ambush rather than the chase. Bears can stand on their hind feet and sit up straight with remarkable balance. Their front paws are flexible enough to grasp fruit and leaves. Bears' non-retractable claws are used for digging, climbing, tearing, and catching prey. The claws on the front feet are larger than those on the back and may be a hindrance when climbing trees; black bears are the most arboreal of the bears, and have the shortest claws. Pandas are unique in having a bony extension on the wrist of the front feet which acts as a thumb, and is used for gripping bamboo shoots as the animals feed.Most mammals have agouti hair, with each individual hair shaft having bands of colour corresponding to two different types of melanin pigment. Bears however have a single type of melanin and the hairs have a single colour throughout their length, apart from the tip which is sometimes a different shade. The coat consists of long guard hairs, which form a protective shaggy covering, and short dense hairs which form an insulating layer trapping air close to the skin. The shaggy coat helps maintain body heat during winter hibernation and is shed in the spring leaving a shorter summer coat. Polar bears have hollow, translucent guard hairs which gain heat from the sun and conduct it to the dark-coloured skin below. They have a thick layer of blubber for extra insulation, and the soles of their feet have a dense pad of fur. Other than the bold black-and-white pelage of the panda, bears tend to be uniform in colour, although some species may have markings on the chest or face.Bears have small rounded ears so as to minimise heat loss, but neither their hearing or sight are particularly acute. Unlike many other carnivorans they have colour vision, perhaps to help them distinguish ripe nuts and fruits. They are unique among carnivorans in not having touch-sensitive whiskers on the muzzle; however, they have an excellent sense of smell, better than that of the dog, or possibly any other mammal. They use smell for signalling to each other (either to warn off rivals or detect mates) and for finding food. Smell is the principal sense used by bears to locate most of their food, and they have excellent memories which helps them to relocate places where they have found food before.(File:Ursus arctos 01 MWNH 145 (cropped).JPG|thumb|left|Brown bear skull)The skulls of bears are massive, providing anchorage for the powerful masseter and temporal jaw muscles. The canine teeth are large but mostly used for display, and the molar teeth flat and crushing. Unlike most other members of the Carnivora, bears have relatively undeveloped carnassial teeth, and their teeth are adapted for a diet that includes a significant amount of vegetable matter.BOOK, Anon, Mammal Anatomy: An Illustrated Guide,weblink 2010, Marshall Cavendish, 978-0-7614-7882-9, 104–123, Considerable variation occurs in dental formula even within a given species. This may indicate bears are still in the process of evolving from a mainly meat-eating diet to a predominantly herbivorous one. Polar bears appear to have secondarily re-evolved carnassial-like cheek teeth, as their diets have switched back towards carnivory.BOOK, Macdonald, D., Bunnell, Fred, 1984, The Encyclopedia of Mammals, Facts on File, 87, 978-0-87196-871-5,weblink Sloth bears lack lower central incisors and use their protusible lips for sucking up the termites on which they feed. The general dental formula for living bears is:{{DentalFormula|upper=3.1.2–4.2|lower=3.1.2–4.3}}. The structure of the larynx of bears appears to be the most basal of the caniforms. They possess air pouches connected to the pharynx which may amplify their vocalisations.JOURNAL, Weissengruber, G. E., Forstenpointner, G., Kübber-Heiss, A., Riedelberger, K., Schwammer, H., Ganzberger, K., 2001, Occurrence and structure of epipharyngeal pouches in bears (Ursidae), Journal of Anatomy, 198, 3, 309–14, 11322723, 1468220,weblink 10.1046/j.1469-7580.2001.19830309.x, 2017-12-12,weblink" title="web.archive.org/web/20171213043736weblink">weblink 2017-12-13, live, Bears have a fairly simple digestive system typical for carnivorans, with a single stomach, short undifferentiated intestines and no cecum.BOOK, Schwartz, C. C., Miller, S. D., Haroldson, M. A., Grizzly Bear, Wild Mammals of North America; biology, management and conservation, Johns Hopkins University Press, Feldhamer, G., Thompson, B., Chapman, J., 562, 978-0-8018-7416-1,weblink 2003-11-19, Even the herbivorous giant panda still has the digestive system of a carnivore, as well as carnivore-specific genes. Its ability to digest cellulose is ascribed to the microbes in its gut.JOURNAL, Ruiqiang Li, The sequence and de novo assembly of the giant panda genome, Nature, 463, 21, 311–317, 2010, 20010809, 3951497, 10.1038/nature08696, harv, 2010Natur.463..311L, Fan, Wei, Tian, Geng, Zhu, Hongmei, He, Lin, Cai, Jing, Huang, Quanfei, Cai, Qingle, Li, Bo, Bai, Yinqi, Zhang, Zhihe, Zhang, Yaping, Wang, Wen, Li, Jun, Wei, Fuwen, Li, Heng, Jian, Min, Li, Jianwen, Zhang, Zhaolei, Nielsen, Rasmus, Li, Dawei, Gu, Wanjun, Yang, Zhentao, Xuan, Zhaoling, Ryder, Oliver A., Leung, Frederick Chi-Ching, Zhou, Yan, Cao, Jianjun, Sun, Xiao, Fu, Yonggui, 29, Bears must spend much of their time feeding in order to gain enough nutrition from foliage. The panda, in particular, spends 12–15 hours a day feeding.

Distribution and habitat

File:Spectacled Bear 161 (2).jpg|thumb|right|The spectacled bearspectacled bear{{Further|List of carnivorans by population}}Extant bears are found in sixty countries primarily in the Northern Hemisphere and are concentrated in Asia, North America, and Europe. An exception is the spectacled bear; native to South America, it inhabits the Andean region. The sun bear's range extends below the equator in Southeast Asia.Ward and Kynaston, p.52 The Atlas bear, a subspecies of the brown bear was distributed in North Africa from Morocco to Libya, but it became extinct around the 1870s.JOURNAL, Hamdine, Watik, Thévenot, Michel, Michaux, Jacques, 1998, Histoire récente de l'ours brun au Maghreb, Comptes Rendus de l'Académie des Sciences, 321, 7, 565–570, 10.1016/S0764-4469(98)80458-7, French, 1998CRASG.321..565H, The most widespread species is the brown bear, which occurs from Western Europe eastwards through Asia to the western areas of North America. The American black bear is restricted to North America, and the polar bear is restricted to the Arctic Sea. All the remaining species of bear are Asian.BOOK, Servheen, C., Herrero, S., Peyton, B., Bears: Status Survey and Conservation Action Plan,weblink 1999, IUCN, 978-2-8317-0462-3, 5–10, They are opportunistic omnivores whose diet varies from plant foliage, roots, and fruits; insect adults, larvae, and eggs; animal matter from carrion; animal matter from predation; and fish. Their dentition and digestive system reflects this varied diet., They occur in a range of habitats which include tropical lowland rainforest, both coniferous and broadleaf forests, prairies, steppes, montane grassland, alpine scree slopes, Arctic tundra and in the case of the polar bear, ice floes. Bears may dig their dens in hillsides or use caves, hollow logs and dense vegetation for shelter.

Behaviour and life history

File:Bear tracks (5062843250).jpg|thumb|right|American black bear tracks at Superior National Forest, MinnesotaMinnesotaBrown and American black bears are generally diurnal, meaning that they are active for the most part during the day, though they may forage substantially by night.JOURNAL, Klinka, D. R., Reimchen, T. E., Nocturnal and diurnal foraging behaviour of brown bears ( Ursus arctos ) on a salmon stream in coastal British Columbia,weblink Canadian Journal of Zoology, 80, 8, 1317–1322, 2002, 10.1139/Z02-123, 2017-01-07,weblink" title="web.archive.org/web/20170812163506weblink">weblink 2017-08-12, live, Other species may be nocturnal, active at night, though female sloth bears with cubs may feed more at daytime to avoid competition from conspecifics and nocturnal predators.Ward and Kynaston, 99 Bears are overwhelmingly solitary and are considered to be the most asocial of all the Carnivora. The only times bears are encountered in small groups are mothers with young or occasional seasonal bounties of rich food (such as salmon runs).BOOK, Sandell, M., 1989, The mating tactics and spacing patterns of solitary carnivores, Carnivore behavior, ecology, and evolution, 164–182, Springer, 978-1-4613-0855-3, 10.1007/978-1-4613-0855-3_7, Fights between males can occur and older individuals may have extensive scarring, which suggests that maintaining dominance can be intense.Ward and Kynaston, p. 130 With their acute sense of smell, bears can locate carcasses from several kilometres away. They use olfaction to locate other foods, encounter mates, avoid rivals and recognise their cubs.

Feeding

File:Giant Panda Tai Shan.JPG|thumb|left|Giant panda feeding on bamboo at Smithsonian National Zoological Park, Washington, D. C. This species is almost entirely herbivorousherbivorousMost bears are opportunistic omnivores and consume more plant than animal matter. They eat anything from leaves, roots, and berries to insects, carrion, fresh meat, and fish, and have digestive systems and teeth adapted to such a diet. At the extremes are the almost entirely herbivorous giant panda and the mostly carnivorous polar bear. However, all bears feed on any food source that becomes seasonally available.Ward and Kynaston, p. 83 For example, Asiatic black bears in Taiwan consume large numbers of acorns when these are most common, and switch to ungulates at other times of the year.JOURNAL, Hwang, Mei-Hsiu, 2002, Diets of Asiatic black bears in Taiwan, with Methodological and Geographical Comparisons,weblink Ursus, 13, 111–125, 2012-12-22,weblink" title="web.archive.org/web/20130723060824weblink">weblink 2013-07-23, live, When foraging for plants, bears choose to eat them at the stage when they are at their most nutritious and digestible, typically avoiding older grasses, sedges and leaves.WEB, Food Habits of Grizzly Bears and Black Bears in the Yellowstone Ecoystem,weblink National Park Service, 18 January 2017,weblink 2017-01-28, live, Hence, in more northern temperate areas, browsing and grazing is more common early in spring and later becomes more restricted. Knowing when plants are ripe for eating is a learned behaviour. Berries may be foraged in bushes or at the tops of trees, and bears try to maximize the number of berries consumed versus foliage. In autumn, some bear species forage large amounts of naturally fermented fruits, which affects their behaviour.BOOK, Rue, Leonard Lee, Furbearing Animals of North America,weblink 1981, Crown Publishers, 978-0-517-53942-2, 129, Smaller bears climb trees to obtain mast (edible reproductive parts, such as acorns).JOURNAL, Mattson, David J., 1998, Diet and Morphology of Extant and Recently Extinct Northern Bears, Ursus, A Selection of Papers from the Tenth International Conference on Bear Research and Management, Fairbanks, Alaska, July 1995, and Mora, Sweden, September 1995, 10, 479–496, 3873160, Such masts can be very important to the diets of these species, and mast failures may result in long-range movements by bears looking for alternative food sources.JOURNAL, Ryan, Christopher, 2007, Influence of mast production on black bear non-hunting mortalities in West Virginia, Ursus, 18, 1, 46–53, 10.2192/1537-6176(2007)18[46:IOMPOB]2.0.CO;2, Pack, James C., Igo, William K., Billings, Anthony, Brown bears, with their powerful digging abilities, commonly eat roots. The panda's diet is over 99% bamboo,BOOK, Lumpkin, Susan, Seidensticker, John, Giant Pandas, Collins, 2007, 978-0-06-120578-1, 63,weblink of 30 different species. Its strong jaws are adapted for crushing the tough stems of these plants, though they prefer to eat the more nutritious leaves.Ward and Kynaston, pp. 89–92WEB,weblink Progress in the utilization of urea-ammonia treated crop residues: biological and socio-economic aspects of animal production and application of the technology on small farms, Dolberg, Frands, 1 August 1992, University of Arhus, 10 August 2010,weblink" title="web.archive.org/web/20110707053325weblink">weblink 2011-07-07, live, Bromeliads can make up to 50% of the diet of the spectacled bear, which also has strong jaws to bite them open.Ward and Kynaston, p. 87File:Bear Alaska (3).jpg|thumb|Brown bear feeding on infrequent, but predictable, salmon migrations in AlaskaAlaskaThe sloth bear, though not as specialised as polar bears and the panda, has lost several front teeth usually seen in bears, and developed a long, suctioning tongue to feed on the ants, termites, and other burrowing insects they favour. At certain times of the year, these insects can make up 90% of their diets.JOURNAL, Joshi, Anup, 1997, Seasonal and Habitat-Related Diets of Sloth Bears in Nepal, Journal of Mammalogy, 1978, 2, 584–597, 10.2307/1382910, Garshelis, David L., Smith, James L. D., 1382910, Some species may raid the nests of wasps and bees for the honey and immature insects, in spite of stinging from the adults.WEB, What do bears like to eat in a beehive?, North American Bear Center,weblink 5 April 2017,weblink 2017-04-06, dead, Sun bears use their long tongues to lick up both insects and honey.Ward and Kynaston, p. 89 Fish are an important source of food for some species, and brown bears in particular gather in large numbers at salmon runs. Typically, a bear plunges into the water and seizes a fish with its jaws or front paws. The preferred parts to eat are the brain and eggs. Small burrowing mammals like rodents may be dug out and eaten.File:Polar bear (Ursus maritimus) with its prey.jpg|thumb|left|Polar bear feeding on a seal on an ice floe north of SvalbardSvalbardThe brown bear and both species of black bears sometimes take large ungulates, such as deer and bovids, mostly the young and weak.JOURNAL, Zager, Peter, Beecham, John, 2006, The role of American black bears and brown bears as predators on ungulates in North America, Ursus, 17, 2, 95–108, 10.2192/1537-6176(2006)17[95:TROABB]2.0.CO;2, Ward and Kynaston, pp. 93–98 These animals may be taken by a short rush and ambush, though hiding young may be stiffed out and pounced on.WEB, Mattson, David, Southwest Biological Science Center, Foraging Behavior of North American Bears,weblinkweblink" title="web.archive.org/web/20070711074612weblink">weblink dead, 11 July 2007, WEB, Animal Protein, North American bear Center,weblink 22 March 2017,weblink 2017-03-23, dead, The polar bear mainly preys on seals, stalking them from the ice or breaking into their dens. They primarily eat the highly digestible blubber.Ward and Kynaston, p. 92 Large mammalian prey is typically killed by a bite to the head or neck, or (in the case of young) simply pinned down and mauled.JOURNAL, French, S. P., French, M. G., Predatory behavior of grizzly bears feeding on elk calves in Yellowstone National Park, 1986–1988, 1990, International Conf. Bear Res. And Manage, 8, 335–341,weblink 10.2307/3872937, 3872937, 2017-04-04,weblink" title="web.archive.org/web/20130722090642weblink">weblink 2013-07-22, live, Predatory behaviour in bears is typically taught to the young by the mother.Bears are prolific scavengers and kleptoparasites, stealing food caches from rodents, and carcasses from other predators.WEB, COSEWIC. Canadian Wildlife Service, Assessment and Update Status Report on the Grizzly Bear (Ursus arctos), Environment Canada, 2002,weblink 8 April 2007, For hibernating species, weight gain is important as it provides nourishment during winter dormancy. A brown bear can eat {{convert|41|kg|lb|abbr=on}} of food and gain {{convert|2–3|kg|lb|abbr=on}} of fat a day prior to entering its den.Ward and Kynaston, p. 104

Communication

File:Ursus thibetanus 01.JPG|thumb|left|Captive Asian black bearAsian black bearBears produce a number of vocal and non-vocal sounds. Tongue-clicking, grunting or chuffing many be made in cordial situations, such as between mothers and cubs or courting couples, while moaning, huffing, sorting or blowing air is made when an individual is stressed. Barking is produced during times of alarm, excitement or to give away the animal's position. Warning sounds include jaw-clicking and lip-popping, while teeth-chatters, bellows, growls, roars and pulsing sounds are made in aggressive encounters. Cubs may squeal, bawl, bleat or scream when in distress and make motor-like humming when comfortable or nursing.Ward and Kynaston, pp. 117–121BOOK, Naughton, D., 2014, The Natural History of Canadian Mammals: Opossums and Carnivores, University of Toronto Press, 218–219, 236, 251–252, 978-1-4426-4483-0, WEB, Vocalizations and Body Language,weblink North American Bear Center, 7 January 2017,weblink 2016-12-23, dead, BOOK, Masterson, Linda, 2016, Living With Bears Handbook: Expanded 2nd Edition, PixyJack Press, 215–16, 978-1-936555-61-1, JOURNAL, Peters, G., Owen, M., Rogers, L., 2007, Humming in bears: a peculiar sustained mammalian vocalization, Acta Theriologica, 52, 4, 379–389, 10.1007/BF03194236,weblink 2017-09-08,weblink" title="web.archive.org/web/20140203154521weblink">weblink 2014-02-03, live, JOURNAL, Laurie, A., Seidensticker, J., 1977, Behavioural ecology of the Sloth bear (Melursus ursinus), Journal of Zoology, 182, 2, 187–204, 10.1111/j.1469-7998.1977.tb04155.x,weblink 2017-03-26,weblink 2017-03-27, live, File:Standing Sloth Bear.jpg|thumb|upright|Sloth bear rubbing against tree at Nagarhole Tiger ReserveNagarhole Tiger ReserveBears sometimes communicate with visual displays such as standing upright, which exaggerates the individual's size. The chest markings of some species may add to this intimidating display. Staring is an aggressive act and the facial markings of spectacled bears and giant pandas may help draw attention to the eyes during agonistic encounters.Ward and Kynaston, pp. 124–125 Individuals may approach each other by stiff-legged walking with the head lowered. Dominance between bears is asserted by making a frontal orientation, showing the canine teeth, muzzle twisting and neck stretching. A subordinate may respond with a lateral orientation, by turning away and dropping the head and by sitting or lying down.JOURNAL, Stonorov, D., Stokes, A. W., 1972, Social behavior of the Alaska brown bear, 2, 232–242,weblink 10.2307/3872587, Bears: Their Biology and Management, 2017-01-11,weblink" title="web.archive.org/web/20130722184419weblink">weblink 2013-07-22, live, 3872587, JOURNAL, Jordan, R. H., 1976, Threat behavior of the black bear (Ursus americanus), Bears: Their Biology and Management, 40, 57–63, 10.2307/3872754, 3872754, Bears may mark territory by rubbing against trees and other objects which may serve to spread their scent. This is usually accompanied by clawing and biting the object. Bark may be spread around to draw attention to the marking post.Ward and Kynaston, p. 122 Pandas are known to mark objects with urine and a waxy substance from their anal glands.JOURNAL, Nie, Y., Swaisgood, R. R., Zhang, Z., Hu, Y., Ma, Y., Wei, F., 2012, Giant panda scent-marking strategies in the wild: role of season, sex and marking surface, Animal Behaviour, 84, 1, 39–44, 10.1016/j.anbehav.2012.03.026, Polar bears leave behind their scent in their tracks which allow individuals to keep track of one another in the vast Arctic wilderness.JOURNAL, Owen, M. A., Swaisgood, R. R., Slocomb, C., Amstrup, S. C., Durner, G. M., Simac, K., Pessier, A. P., 2014, An experimental investigation of chemical communication in the polar bear, Journal of Zoology, 295, 1, 36–43, 10.1111/jzo.12181,

Reproduction and development{{anchor|Breeding}}

File:Black Bears mating.jpg|thumb|left|American black bears mating at the North American Bear CenterNorth American Bear CenterThe mating system of bears has variously been described as a form of polygyny, promiscuity and serial monogamy.WEB, Eide, S., Miller, S., Brown Bear, Alaska Department of Fish and Game,weblink 14 January 2016,weblink" title="web.archive.org/web/20130303124834weblink">weblink 2013-03-03, live, JOURNAL, Lariviere, S., 2001, Ursus americanus, Mammalian Species, 647, 1–11,weblink 10.1644/1545-1410(2001)6472.0.co;2, dead,weblink" title="web.archive.org/web/20160316025524weblink">weblink 2016-03-16, During the breeding season, males take notice of females in their vicinity and females become more tolerant of males. A male bear may visit a female continuously over a period of several days or weeks, depending on the species, to test her reproductive state. During this time period, males try to prevent rivals from interacting with their mate. Courtship may be brief, although in some Asian species, courting pairs may engage in wrestling, hugging, mock fighting and vocalising. Ovulation is induced by mating, which can last up to 30 minutes depending on the species.Ward and Kynaston, pp. 138–141File:Cub polar bear is nursing 2.OGG|thumb|Polar bearPolar bearGestation typically lasts 6–9 months, including delayed implantation, and litter size numbers up to four cubs. Giant pandas may give birth to twins but they can only suckle one young and the other is left to die.WEB,weblink Panda Facts, Pandas International, 26 August 2015, dead,weblink" title="web.archive.org/web/20150924063658weblink">weblink 24 September 2015, In northern living species, birth takes place during winter dormancy. Cubs are born blind and helpless with at most a thin layer of hair, relying on their mother for warmth. The milk of the female bear is rich in fat and antibodies and cubs may suckle for up to a year after they are born. By 2–3 months, cubs can follow their mother outside the den. They usually follow her on foot, but sloth bear cubs may ride on their mother's back.Ward and Kynaston, pp. 144–148BOOK, Nowak, R. M., 2005, Walker's Carnivores of the World, Johns Hopkins University Press, 114, 978-0-8018-8032-2, Male bears play no role in raising young. Infanticide, where an adult male kills the cubs of another, has been recorded in polar bears, brown bears and American black bears but not in other species.Ward and Kynaston, p. 132 Males kill young to bring the female into oestrus.JOURNAL, 10.1111/j.1439-0310.2006.01152.x, Mating Strategies in Relation to Sexually Selected Infanticide in a Non-Social Carnivore: The Brown Bear,weblink Ethology, 112, 3, 238–246, 2006, Bellemain, Eva, Swenson, Jon E., Taberlet, Pierre, 2017-05-02,weblink" title="web.archive.org/web/20161221015155weblink">weblink 2016-12-21, live, Cubs may flee and the mother defends them even at the cost of her life.JOURNAL, Bellemain, E., Zedrosser, A., Manel, S., Waits, L. P., Taberlet, P., Swenson, J. E., 2005, 16543170, 1560043, The dilemma of female mate selection in the brown bear, a species with sexually selected infanticide, Proceedings of the Royal Society of London B: Biological Sciences, 273, 1584, 283–291, 10.1098/rspb.2005.3331, JOURNAL, Swenson, J. E., Dahle, B., Sandegren, F., 2001, Intraspecific predation in Scandinavian brown bears older than cubs-of-the-year, Ursus, 12, 81–91, 3873233,weblink 2017-01-14,weblink 2017-11-16, live, JOURNAL, Mörner, T., Eriksson, H., Bröjer, C., Nilsson, K., Uhlhorn, H., Ågren, E., af Segerstad, C.H., Jansson, D.S., Gavier-Widén, D., 2005, Diseases and mortality in free-ranging brown bear (Ursus arctos), gray wolf (Canis lupus), and wolverine (Gulo gulo) in Sweden, Journal of Wildlife Diseases, 41, 2, 298–303, 16107663, 10.7589/0090-3558-41.2.298, In some species, offspring may become independent around the next spring, through some may stay until the female successfully mates again. Bears reach sexual maturity shortly after they disperse; at around 3–6 years depending on the species. Male Alaskan brown bears and polar bears may continue to grow until they are 11 years old. Lifespan may also vary between species. The brown bear can live an average of 25 years.WEB, Grizzly Bear, National Geographic.com, 29 April 2017,weblinkweblink" title="web.archive.org/web/20170421214527weblink">weblink 2017-04-21, live,

Hibernation{{anchor|Winter_dormancy}}

Bears of northern regions, including the American black bear and the grizzly bear, hibernate in the winter.JOURNAL, Gerhard Heldmeier, 2011, Life on low flame in hibernation, Science, 331, 6019, 866–867, 10.1126/science.1203192, 21330523, 2011Sci...331..866H, JOURNAL, Shimozuru, M., 2013, Pregnancy during hibernation in Japanese black bears: effects on body temperature and blood biochemical profiles, Journal of Mammalogy, 94, 3, 618–627, 10.1644/12-MAMM-A-246.1, etal, During hibernation, the bear's metabolism slows down, its body temperature decreases slightly, and its heart rate slows from a normal value of 55 to just 9 beats per minute.JOURNAL, Tøien, Ø., 2011, Hibernation in Black Bears: Independence of Metabolic Suppression from Body Temperature, Science, 331, 6019, 906–909, 10.1126/science.1199435, 21330544, etal, 2011Sci...331..906T, Bears normally do not wake during their hibernation, and can go the entire period without eating, drinking, urinating, or defecating. A fecal plug is formed in the colon, and is expelled when the bear wakes in the spring.WEB, Biel, M, Gunther, K, Denning and Hibernation Behavior,weblink Yellowstone National Park, National Park Service, 18 November 2016,weblink 2016-11-18, live, If they have stored enough body fat, their muscles remain in good condition, and their protein maintenance requirements are met from recycling waste urea. Female bears give birth during the hibernation period, and are roused when doing so.

Mortality

File:Björnjakt i Dalarna - Nordiska Museet - NMA.0052736.jpg|thumb|right|Hunters with shot bear, Sweden, early 20th century. This photograph is in the Nordic MuseumNordic MuseumBears do not have many predators. The most important are humans, and as they started cultivating crops, they increasingly came in conflict with the bears that raided them. Since the invention of firearms, people have been able to kill bears with greater ease.JOURNAL, McTaggart Cowan, I., 1972, The Status and Conservation of Bears (Ursidae) of the World: 1970, Bears: Their Biology and Management, 2, 343–367, 10.2307/3872596, 10.1.1.483.1402, 3872596, Felids like the tiger may also prey on bears,BOOK,weblink The ecology, behavior, management and conservation status of brown bears in Sikhote-Alin, ru, Seryodkin, Ivan, 2006, Far Eastern National University, Vladivostok, Russia, 1–252, dead,weblink" title="web.archive.org/web/20131224090426weblink">weblink 2013-12-24, JOURNAL,weblink Seryodkin, 2003, Denning ecology of brown bears and Asiatic black bears in the Russian Far East, Ursus, 14, 2, 159, etal, 2014-10-05,weblink" title="web.archive.org/web/20110817151324weblink">weblink 2011-08-17, live, particularly cubs, which may also be threatened by canids.Bears are parasitized by eighty species of parasites, including single-celled protozoans and gastro-intestinal worms, and nematodes and flukes in their heart, liver, lungs and bloodstream. Externally they have ticks, fleas and lice. A study of American black bears found seventeen species of endoparasite including the protozoan Sarcocystis, the parasitic worm Diphyllobothrium mansonoides, and the nematodes Dirofilaria immitis, Capillaria aerophila, Physaloptera sp., Strongyloides sp. and others. Of these, D. mansonoides and adult C. aerophila were causing pathological symptoms.JOURNAL, Crum, James M., Nettles, Victor F., Davidson, William R., 1978, Studies on endoparasites of the black bear (Ursus americanus) in the southeastern United States, Journal of Wildlife Diseases, 14, 2, 178–186, 10.7589/0090-3558-14.2.178, By contrast, polar bears have few parasites; many parasitic species need a secondary, usually terrestrial, host, and the polar bear's life style is such that few alternative hosts exist in their environment. The protozoan Toxoplasma gondii has been found in polar bears, and the nematode Trichinella nativa can cause a serious infection and decline in older polar bears.BOOK, Derocher, Andrew E., Polar Bears: A Complete Guide to Their Biology and Behavior,weblink 2012, JHU Press, 978-1-4214-0305-2, 212, Bears in North America are sometimes infected by a Morbillivirus similar to the canine distemper virus.BOOK, Emergence and Control of Zoonotic Ortho- and Paramyxovirus Diseases,weblink John Libbey Eurotext, 978-2-7420-0392-1, 167, They are susceptible to infectious canine hepatitis (CAV-1), with free-living black bears dying rapidly of encephalitis and hepatitis.BOOK, Williams, Elizabeth S., Barker, Ian K., Infectious Diseases of Wild Mammals,weblink 2008, John Wiley & Sons, 978-0-470-34481-1, 203,

Relationship with humans

Conservation

{{multiple image |align=right |direction=vertical |
caption1=Giant pandas at the Sichuan Giant Panda Sanctuaries caption2=A barrel trap in Grand Teton National Park, Wyoming used to relocate bears away from where they might attack humans}}In modern times, bears have come under pressure through encroachment on their habitatsWEB, Brown Bear – Threats Grizzlies: Found in 2% of their former range,weblink WWF, 16 January 2017,weblink" title="web.archive.org/web/20161221195824weblink">weblink 2016-12-21, live, and illegal trade in bear parts, including the Asian bile bear market, though hunting is now banned, largely replaced by farming.WEB, Bacon, Heather, Implications of bear bile farming,weblink Vet Times, 16 January 2017, 12 May 2008,weblink 2017-01-18, live, The IUCN lists six bear species as vulnerable;WEB, Keyword search: "Ursidae", Exact phrase, The entire database,weblink IUCN, 16 January 2017,weblink" title="web.archive.org/web/20110805065610weblink">weblink 2011-08-05, live, even the two least concern species, the brown bear and the American black bear, are at risk of extirpation in certain areas. In general these two species inhabit remote areas with little interaction with humans, and the main non-natural causes of mortality are hunting, trapping, road-kill and depredation.BOOK,weblink Chapter 8. American Black Bear Conservation Action Plan, Pelton, Michael R., Coley, Alex B., Eason, Thomas H., Doan Martinez, Diana L., Pederson, Joel A., van Manen, Frank T., Weaver, Keith M., 1999, IUCN, 978-2-8317-0462-3, 144–156, Laws have been passed in many areas of the world to protect bears from habitat destruction. Public perception of bears is often positive, as people identify with bears due to their omnivorous diets, their ability to stand on two legs, and their symbolic importance.JOURNAL, Kellert, Stephen, Public Attitudes toward Bears and Their Conservation, Bears: Their Biology and Management, 9, 1, 43–50, 10.2307/3872683, 1994, 3872683, Support for bear protection is widespread, at least in more affluent societies.JOURNAL, Andersone, Žanete, 2004, Public perception of large carnivores in Latvia, Ursus, 15, 2, 181–187, 10.2192/1537-6176(2004)0152.0.CO;2, Ozolinš, Jānis, The giant panda has become a worldwide symbol of conservation. The Sichuan Giant Panda Sanctuaries, which are home to around 30% of the wild panda population, gained a UNESCO World Heritage Site designation in 2006.WEB,weblink Pandas gain world heritage status, BBC News, 2019-08-22, Where bears raid crops or attack livestock, they may come into conflict with humans.JOURNAL, Goldstein, Isaac, 2006, Andean bear–livestock conflicts: a review, Ursus, 17, 1, 8–15, 10.2192/1537-6176(2006)17[8:ABCAR]2.0.CO;2, Paisley, Susanna, Wallace, Robert, Jorgenson, Jeffrey P., Cuesta, Francisc, Castellanos, Armando, JOURNAL, Fredriksson, Gabriella, 2005, Human–sun bear conflicts in East Kalimantan, Indonesian Borneo, Ursus, 16, 1, 130–137, 10.2192/1537-6176(2005)016[0130:HBCIEK]2.0.CO;2, In poorer rural regions, attitudes may be more shaped by the dangers posed by bears, and the economic costs they cause to farmers and ranchers.

Attacks

Several bear species are dangerous to humans, especially in areas where they have become used to people; elsewhere, they generally avoid humans. Injuries caused by bears are rare, but are widely reported.JOURNAL, Clark, Douglas, 2003, Polar Bear–Human Interactions in Canadian National Parks, 1986–2000,weblink Ursus, 14, 1, 65–71, 2012-12-22,weblink" title="web.archive.org/web/20130722191859weblink">weblink 2013-07-22, live, Bears may attack humans in response to being startled, in defense of young or food, or even for predatory reasons.WEB, Than, K., 2013, Maulings by Bears: What's Behind the Recent Attacks?, National Geographic.com, 16 January 2017,weblinkweblink" title="web.archive.org/web/20170128084840weblink">weblink 2017-01-28, live,

Entertainment, hunting, food and folk medicine

File:The dancing bear by William Frederick Witherington.jpg|thumb|left|The dancing bear by William Frederick WitheringtonWilliam Frederick WitheringtonBears in captivity have for centuries been used for entertainment. They have been trained to dance,BOOK, Findeizen, Nikolai, History of Music in Russia from Antiquity to 1800, Vol. 1: From Antiquity to the Beginning of the Eighteenth Century,weblink 2008, Indiana University Press, 978-0-253-02637-8, 201, and were kept for baiting in Europe at least since the 16th century. There were five bear-baiting gardens in Southwark, London at that time; archaeological remains of three of these have survived.WEB, Elizabethan Playhouses and Bear Baiting Arenas Given Protection,weblink Historic England, 4 January 2017, 26 September 2016,weblink 2017-01-04, live, Across Europe, nomadic Romani bear handlers called Ursari lived by busking with their bears from the 12th century.BOOK, Fraser, Angus M., The Gypsies, Blackwell, 1995, 45–48, 226, 978-0-631-19605-1, File:Theodor Aman - Ursarul.jpg|thumb|A nomadic ursar, a Romani bear-busker. Drawing by Theodor AmanTheodor AmanBears have been hunted for sport, food, and folk medicine. Their meat is dark and stringy, like a tough cut of beef. In Cantonese cuisine, bear paws are considered a delicacy. Bear meat should be cooked thoroughly, as it can be infected with the parasite Trichinella spiralis.WEB,weblink 4 October 2006, Trichinellosis Associated with Bear Meat,weblink" title="web.archive.org/web/20060930201027weblink">weblink 30 September 2006, live, NEWS,weblink 4 October 2006, Bear meat poisoning in Siberia, BBC News, 21 December 1997,weblink" title="web.archive.org/web/20081211191933weblink">weblink 2008-12-11, live, The peoples of eastern Asia use bears' body parts and secretions (notably their gallbladders and bile) as part of traditional Chinese medicine. More than 12,000 bears are thought to be kept on farms in China, Vietnam, and South Korea for the production of bile. Trade in bear products is prohibited under CITES, but bear bile has been detected in shampoos, wine and herbal medicines sold in Canada, the United States and Australia.NEWS, Richard, Black,weblink BBC Test kit targets cruel bear trade, BBC News, 11 June 2007, 1 January 2010,weblink" title="web.archive.org/web/20090115175324weblink">weblink 15 January 2009, live,

Literature, art and symbolism

{{further|Bear worship|Jean de l'Ours|Berserker|Kalevala}}{{multiple image |align=right |direction=vertical |width=200
|image1=ShunsenOniguma.jpg |caption1=Onikuma, a Japanese demon bear from Ehon Hyaku Monogatari, c. 1841
|image2=Brīvības_piemineklis-Lāčplēsis.png
|caption2=The Latvian legendary hero Lāčplēsis kills a bear with his bare hands
}}There is evidence of prehistoric bear worship, though this is disputed by archaeologists.JOURNAL, Wunn, Ina, 2000, Beginning of Religion, Numen (journal), Numen, 47, 4, 417–452, 10.1163/156852700511612, It is possible that bear worship existed in early Chinese and Ainu cultures.JOURNAL, The Concepts behind the Ainu Bear Festival (Kumamatsuri), Kyōsuke, Kindaichi, Minori, Yoshida, Southwestern Journal of Anthropology, 5, 4, Winter 1949, 345–350, 3628594, 10.1086/soutjanth.5.4.3628594, The prehistoric Finns,JOURNAL, Bonser, Wilfrid, Folklore, The mythology of the Kalevala, with notes on bear-worship among the Finns, 39, 4, 1928, 344–358, 1255969, 10.1080/0015587x.1928.9716794, Siberian peoplesBOOK, Chaussonnet, Valerie, 1995, Native Cultures of Alaska and Siberia, Arctic Studies Center, Washington, D.C., 112, 978-1-56098-661-4, and more recently Koreans considered the bear as the spirit of their forefathers.BOOK, Lee, Jung Young, Korean Shamanistic Rituals, Mouton De Gruyter, 1981, 978-90-279-3378-2, 14, 20, In many Native American cultures, the bear is a symbol of rebirth because of its hibernation and re-emergence.Ward and Kynaston, p. 17 The image of the mother bear was prevalent throughout societies in North America and Eurasia, based on the female's devotion and protection of her cubs.Ward and Kynaston, pp. 12–13 Japanese folklore features the Onikuma, a "demon bear" that walks upright.WEB, Davisson, Zack, Onikuma – Demon Bear,weblink Hyakumonogatari Kaidankai, 19 February 2017, 28 May 2013,weblink 2017-02-20, live, The Ainu of northern Japan, a different people from the Japanese, saw the bear instead as sacred; Hirasawa Byozan painted a scene in documentary style of a bear sacrifice in an Ainu temple, complete with offerings to the dead animal's spirit.BOOK, Davidson, Peter, The Idea of North,weblink 2005, Reaktion Books, 978-1-86189-230-0, 179, in the Meiji period .. handscroll of paintings of Ainu dwellings and customs .. The painter was Hirasawa Byozan and he titled the work Scenes of the Daily Life of the Ezo. His paintings are documentary, even anthropological in intent, for all their beauty., In Korean mythology, a tiger and a bear prayed to Hwanung, the son of the Lord of Heaven, that they might become human. Upon hearing their prayers, Hwanung gave them 20 cloves of garlic and a bundle of mugwort, ordering them to eat only this sacred food and remain out of the sunlight for 100 days. The tiger gave up after about twenty days and left the cave. However, the bear persevered and was transformed into a woman. The bear and the tiger are said to represent two tribes that sought the favor of the heavenly prince.WEB,weblink The Myth of Gojoseon's Founding-King Dan-gun, 2017-08-29,weblink" title="web.archive.org/web/20170828194504weblink">weblink 2017-08-28, live, The bear-woman (Ungnyeo; 웅녀/) was grateful and made offerings to Hwanung. However, she lacked a husband, and soon became sad and prayed beneath a "divine birch" tree ({{korean|hangul=신단수|hanja=|rr=shindansu}}) to be blessed with a child. Hwanung, moved by her prayers, took her for his wife and soon she gave birth to a son named Dangun Wanggeom – who was the legendary founder of Gojoseon, the first ever Korean kingdom.BOOK, weblink, Tudor, Daniel, 2013, Korea: The Impossible Country: The Impossible Country, Tuttle Publishing, 978-1462910229, File:Sidney Hall - Urania's Mirror - Ursa Major.jpg|thumb|left|The constellation of Ursa Major as depicted in Urania's MirrorUrania's MirrorArtio (Dea Artio in the Gallo-Roman religion) was a Celtic bear goddess. Evidence of her worship has notably been found at Bern, itself named for the bear. Her name is derived from the Celtic word for "bear", artos.BOOK, Room, Adrian,weblink Placenames of the World: Origins and Meanings of the Names for 6,600 Countries, Cities, Territories, Natural Features, and Historic Sites, McFarland, 2006, 57, 9780786422487, In ancient Greece, archaic cult of Artemis in bear form survived into Classical times at Brauron, where young Athenian girls passed an initiation right as arktai "she bears".Burkert, Walter, Greek Religion, 1985:263. For Artemis and one of her nymphs as a she-bear, see the myth of Callisto.The constellations of Ursa Major and Ursa Minor, the great and little bears, are named for their supposed resemblance to bears, from the time of Ptolemy.{{efn|Ptolemy named the constellations in Greek, Ἄρκτος μεγάλη (Arktos Megale) and Ἄρκτος μικρά (Arktos Mikra), the great and little bears.WEB, Ridpath, Ian, Ian Ridpath, Ptolemy's Almagest First printed edition, 1515,weblink 13 January 2017, }} The nearby star Arcturus means "guardian of the bear", as if it were watching the two constellations.WEB,weblink Ἀρκτοῦρος, Liddell, Henry George, Scott, Robert, A Greek-English Lexicon, Perseus, 23 January 2017,weblink" title="web.archive.org/web/20170307140100weblink">weblink 2017-03-07, live, Ursa Major has been associated with a bear for as much as 13,000 years since Paleolithic times, in the widespread Cosmic Hunt myths. These are found on both sides of the Bering land bridge, which was lost to the sea some 11,000 years ago.MAGAZINE, Schaefer, Bradley E., The Origin of the Greek Constellations: Was the Great Bear constellation named before hunter nomads first reached the Americas more than 13,000 years ago?, Scientific American, November 2006, reviewed at WEB, Brown, Miland,weblink The Origin of the Greek Constellations, World History Blog, 30 October 2006, 9 April 2017, ;,weblink" title="web.archive.org/web/20170401164801weblink">weblink 2017-04-01, live, JOURNAL, Berezkin, Yuri, The cosmic hunt: variants of a Siberian – North-American myth, Folklore, 31, 2005, 79–100, 10.7592/FEJF2005.31.berezkin, Pliny the Elder's Natural History (1st century AD) claims that "when first born, [bears] are shapeless masses of white flesh, a little larger than mice; their claws alone being prominent. The mother then licks them gradually into proper shape."BOOK, Pliny,weblink Natural History, 8.4], Bostock, John, Riley, Henry T., 1855, This belief was echoed by authors of bestiaries throughout the medieval period.WEB, Badke, David,weblink The Medieval Bestiary: Bear, 23 January 2017,weblink" title="web.archive.org/web/20170312052954weblink">weblink 2017-03-12, live, File:The Three Bears - Project Gutenberg eText 17034.jpg|thumb|right|upright=0.9|"The Three Bears", Arthur Rackham's illustration to English Fairy Tales, by Flora Annie SteelFlora Annie SteelBears are mentioned in the Bible; the Second Book of Kings relates the story of the prophet Elisha calling on them to eat the youths who taunted him.Second Book of Kings, 2:23–25 Legends of saints taming bears are common in the Alpine zone. In the arms of the bishopric of Freising, the bear is the dangerous totem animal tamed by St. Corbinian and made to carry his civilised baggage over the mountains. Bears similarly feature in the legends of St. Romedius, Saint Gall and Saint Columbanus. This recurrent motif was used by the Church as a symbol of the victory of Christianity over paganism.BOOK, Pastoreau, Michel, 2007, L'ours. Historie d'un roi déchu, Seuil, French, 978-2-02-021542-8, In the Norse settlements of northern England during the 10th century, a type of "hogback" grave cover of a long narrow block of stone, with a shaped apex like the roof beam of a long house, is carved with a muzzled, thus Christianised, bear clasping each gable end, as in the church at Brompton, North Yorkshire and across the British Isles.BOOK, Hall, Richard, 1995, Viking Age Archaeology, 43 and fig. 22, 978-0-7478-0063-7, File:As Between Friends (Punch magazine, 13 December 1911, detail).jpg|thumb|left|upright=0.9|The Persian Cat, British Lion and Russian Bear in the Anglo-Russian EntenteAnglo-Russian EntenteLāčplēsis, meaning "Bear-slayer", is a Latvian legendary hero who is said to have killed a bear by ripping its jaws apart with his bare hands. However, as revealed in the end of the long epic describing his life, Lāčplēsis' own mother had been a she-bear, and his superhuman strength resided in his bear ears. The modern Latvian military award Order of Lāčplēsis, called for the hero, is also known as The Order of the Bear-Slayer.{{citation needed|date=August 2019}}Bears are popular in children's stories, including Winnie the Pooh,WEB,weblink Pooh celebrates his 80th birthday, BBC News, 24 December 2005, 23 January 2017,weblink" title="web.archive.org/web/20060425143300weblink">weblink 2006-04-25, live, Paddington Bear,WEB, Paddington.com,weblink About, 19 January 2017, dead,weblink" title="web.archive.org/web/20160817131744weblink">weblink 17 August 2016, Gentle BenWEB, Walt Morey, 84; Author of 'Gentle Ben', Associated Press,weblink 14 January 1992, 6 April 2017,weblink" title="web.archive.org/web/20161023134546weblink">weblink 2016-10-23, live, and "The Brown Bear of Norway".BOOK, Kennedy, Patrick, The Brown Bear of Norway, Legendary Fictions of the Irish Celts, Macmillan, 1866,weblink 57–67, An early version of "Goldilocks and the Three Bears",JOURNAL, Elms, Alan C., July–September 1977, "The Three Bears": Four Interpretations, The Journal of American Folklore, 90, 357, 257–273, 539519, 10.2307/539519, was published as "The Three Bears" in 1837 by Robert Southey, many times retold, and illustrated in 1918 by Arthur Rackham.BOOK, Ashliman, D. L., D. L. Ashliman, Folk and Fairy Tales: A Handbook,weblink 2004, Greenwood Publishing Group, 978-0-313-32810-7, 114–115, The Hanna-Barbera character Yogi Bear has appeared in numerous comic books, animated television shows and films.BOOK, Mallory, Michael, Hanna-Barbera Cartoons, Hugh Lauter Levin, 1998, 978-0-88363-108-9, 44, BOOK, Browne, Ray B., Browne, Pat, The Guide to United States Popular Culture,weblink 2001, Popular Press, 978-0-87972-821-2, 944, The Care Bears began as greeting cards in 1982, and were featured as toys, on clothing and in film.NEWS, Holmes, Elizabeth, 9 February 2007, Care Bears Receive a (Gentle) Makeover, Wall Street Journal,weblink 27 January 2017,weblink 2018-01-18, live, Around the world, many children—and some adults—have teddy bears, stuffed toys in the form of bears, named after the American statesman Theodore Roosevelt when in 1902 he had refused to shoot an American black bear tied to a tree.WEB, David Cannadine, Cannadine, David,weblink A Point of View: The grownups with teddy bears, 1 February 2013, BBC, 21 January 2017,weblink" title="web.archive.org/web/20170425025929weblink">weblink 2017-04-25, live,
Bears, like other animals, may symbolize nations. In 1911, the British satirical magazine Punch published a cartoon about the Anglo-Russian Entente by Leonard Raven-Hill in which the British lion watches as the Russian bear sits on the tail of the Persian cat.JOURNAL, Raven-Hill, Leonard, Leonard Raven-Hill, As Between Friends, Punch, 13 December 1911, 141, 429,weblink 19 February 2017,weblink" title="web.archive.org/web/20170220015023weblink">weblink 2017-02-20, live, The Russian Bear has been a common national personification for Russia from the 16th century onwards.NEWS,weblink What the West thinks about Russia is not necessarily true, Telegraph, 23 April 2009, 3 January 2017,weblink" title="web.archive.org/web/20151206043929weblink">weblink 2015-12-06, live, Smokey Bear has become a part of American culture since his introduction in 1944, with his message "Only you can prevent forest fires".WEB,weblink Forest Fire Prevention – Smokey Bear (1944–Present), Ad Council, 1944-08-09, 2010-10-16,weblink" title="web.archive.org/web/20101202162355weblink">weblink 2010-12-02, dead, In the United Kingdom, the bear and staff feature on the heraldic arms of the county of Warwickshire.WEB,weblink Civic Heraldry of England and Wales-Warwickshire, 2011-01-06,weblink" title="web.archive.org/web/20110516001919weblink">weblink 2011-05-16, live, Bears appear in the canting arms of two cities, Bern and Berlin.WEB, The first Buddy Bears in Berlin,weblink Buddy Bär Berlin, 30 January 2014, 2008,weblink" title="web.archive.org/web/20140219023945weblink">weblink 2014-02-19, live,

Organisations

File:Baby Pandas.JPG|thumb|right|Juvenile pandas at the ChengduChengduThe International Association for Bear Research & Management, also known as the International Bear Association, and the Bear Specialist Group of the Species Survival Commission, a part of the International Union for Conservation of Nature focus on the natural history, management, and conservation of bears. Bear Trust International works for wild bears and other wildlife through four core program initiatives, namely Conservation Education, Wild Bear Research, Wild Bear Management, and Habitat Conservation.WEB,weblink Vision and Mission, 2002–2012, Bear Trust International, 8 March 2014,weblink" title="web.archive.org/web/20140226195504weblink">weblink 2014-02-26, live, Specialty organisations for each of the eight species of bears worldwide include:
  • Vital Ground, for the brown bearWEB,weblink Vital Ground, 2019-02-05,weblink 2019-01-20, live,
  • Moon Bears, for the Asiatic black bearWEB,weblink Moon Bears, 2014-03-09,weblink" title="web.archive.org/web/20140309053102weblink">weblink 2014-03-09, live,
  • Black Bear Conservation Coalition, for the North American black bearWEB,weblink Black Bear Conservation Coalition, 2014-03-09,weblink" title="web.archive.org/web/20140103020114weblink">weblink 2014-01-03, live,
  • Polar Bears International, for the polar bearWEB,weblink Polar Bears International, 2014-03-09,weblink" title="web.archive.org/web/20140308234451weblink">weblink 2014-03-08, live,
  • Bornean Sun Bear Conservation Centre, for the sun bearWEB,weblink Bornean Sun Bear Conservation Centre, dead,weblink" title="web.archive.org/web/20140218113140weblink">weblink 2014-02-18,
  • Wildlife SOS, for the sloth bearWEB,weblink Wildlife SOS,
  • Andean Bear Conservation Project, for the Andean bearWEB,weblink Andean Bear Conservation Project, 2014-03-09,weblink" title="web.archive.org/web/20140218004303weblink">weblink 2014-02-18, live,
  • Chengdu Research Base of Giant Panda Breeding, for the giant pandaWEB,weblink Chengdu Research Base of Giant Panda Breeding, 2014-03-09,weblink" title="web.archive.org/web/20140315221701weblink">weblink 2014-03-15, live,

See also

Notes

{{notelist}}

References

{{Reflist|30em}}

Bibliography

  • BOOK, Ward, P., Kynaston, S., 1995, Wild Bears of the World, Facts on File, Inc, 978-0-8160-3245-7,

Further reading

  • BOOK, Bears of the World, Terry, Domico, Mark, Newman, Facts on File, Inc., 1988, 978-0-8160-1536-8,
  • BOOK,weblink The Bear, William Faulkner, William, Faulkner, 1942, Curley Publishing, 978-0-7927-0537-6,
  • BOOK, Brunner, Bernd, Bears: A Brief History, Yale University Press, 2007, 978-0-300-12299-2,

External links

{{commons|Ursidae}} {{Carnivora|C.}}{{Living things in culture}}{{Heraldic creatures}}{{Taxonbar|from=Q11788}}{{Authority control}}


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