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| birth_place = Northumberland, UK| nationality = British| death_date = | death_place = | occupation = Paleontologist, Entomologist| work_institution = University of Illinois at Urbana-Champaign 2009–University of Portsmouth 2001–2009| alma_mater = University of Portsmouth Ph.D, 2009; B.Sc. (Hons), 2004| doctoral_advisor = Dr David M. Martill (University of Portsmouth)| academic_advisors = Prof. Dr. Paul A. Selden (University of Kansas)| spouse = Tina Louise Heads}}Sam W. Heads (born 22 April 1983 in Northumberland, UK) is a British palaeontologist, a Fellow of the Linnean Society of London, a Fellow of the Royal Entomological Society, as well as a former Officer and Editor-in-Chief at the Orthopterists' Society.WEB,weblink The Orthopterists' Society, 140.247.119.138, 22 May 2022,weblink" title="web.archive.org/web/20060813151705weblink">weblink 13 August 2006, dead, He received his training at the University of Portsmouth where he was awarded a B.Sc. (Hons) in Palaeobiology and Evolution in 2004 and a Ph.D. in Entomology in 2009. Dr. Heads is an authority on the taxonomy, systematics and palaeontology of Orthoptera.WEB,weblink The Orthopterists' Society, 140.247.119.138, 22 May 2022,weblink" title="web.archive.org/web/20060813151705weblink">weblink 13 August 2006, dead, He is based at the Illinois Natural History Survey, University of Illinois at Urbana-Champaign.WEB,weblink W. Heads--Illinois Natural History Survey, 1 November 2009, 5 March 2010,weblink" title="web.archive.org/web/20100305200429weblink">weblink dead,

Select publications

• Heads, S.W. & Wang, Y. 2013. First fossil record of Melanoplus differentialis (Orthoptera: Acrididae: Melanoplinae). Entomological News 123: 33–37.JOURNAL, Heads, Sam W., Wang, Yinan, May 2013, First Fossil Record of Melanoplus differentialis (Orthoptera: Acrididae: Melanoplinae),weblink Entomological News, 123, 1, 33–37, 10.3157/021.123.0108, 84197628, 0013-872X,
• Heads, S.W., Taylor, S.J. & Krejca, J.K. 2013. First record of Scapteriscus abbreviatus from Belize (Orthoptera: Gryllotalpidae). Entomological News 123: 241–244.JOURNAL, Heads, Sam W., Taylor, Steven J., Krejca, Jean K., October 2013, First Record of Scapteriscus abbreviatus from Belize (Orthoptera: Gryllotalpidae),weblink Entomological News, 123, 3, 241–244, 10.3157/021.123.0311, 86079147, 0013-872X,
• Hollier, J. & Heads, S.W. 2014. The type specimens of Orthoptera described by American entomologists in the Muséum d’histoire naturelle de Genève. Revue suisse de Zoology 121: 63–76.JOURNAL, Hollier, John, Heads, Sam William, March 2014, The type specimens of Orthoptera described by American entomologists in the collection of the Muséum d'histoire naturelle de Genève,weblink Revue Suisse de Zoologie, 121, 1, 63–76, 0035-418X,
• Barling, N., Heads, S.W. & Martill, D.M. 2013. A new parasitoid wasp (Hymenoptera: Chalcidoidea) from the Lower Cretaceous Crato Formation of Brazil: the first Mesozoic Pteromalidae. Cretaceous Research 45: 258–264,JOURNAL, Barling, Nathan, Heads, Sam W., Martill, David M., 2013-10-01, A new parasitoid wasp (Hymenoptera: Chalcidoidea) from the Lower Cretaceous Crato Formation of Brazil: The first Mesozoic Pteromalidae,weblink Cretaceous Research, en, 45, 258–264, 10.1016/j.cretres.2013.05.001, 2013CrRes..45..258B, 0195-6671,
• Hollier, J., Bruckner, H. & Heads, S.W. 2013. An annotated list of the Orthoptera (Insecta) species described by Henri de Saussure, with an account of the primary type material housed in the Muséum d’histoire naturelle de Genève, Part 5: Grylloidea. Revue suisse de Zoology 120: 445–535.WEB, An annotated list of the Orthoptera (Insecta) species described by Henri de Saussure, with an account of the primary type material housed in the Muséum d'histoire naturelle de Genève, Part 5: Grylloidea {{!, Request PDF|url=https://www.researchgate.net/publication/257644739|access-date=2022-01-31|website=ResearchGate|language=en}}
• Jepson, J.E., Heads, S.W., Makarkin, V.N. & Ren, D. 2013. New fossil mantidflies (Insecta, Neuroptera, Mantispidae) from the Mesozoic of northeastern China. Palaeontology 56: 603–613.JOURNAL, Jepson, James E., Heads, Sam W., Makarkin, Vladimir N., Ren, Dong, 2013, New fossil mantidflies (Insecta: Neuroptera: Mantispidae) from the Mesozoic of north-eastern China, Palaeontology, en, 56, 3, 603–613, 10.1111/pala.12005, 2013Palgy..56..603J, 55218976, 1475-4983, free,
• Heads, S.W., Penney, D. & Green, D.I. 2012. A new species of the cricket genus Proanaxipha in Miocene amber from the Dominican Republic (Orthoptera, Gryllidae, Pentacentrinae). ZooKeys 229: 111–118.JOURNAL, Heads, Sam W., Penney, David, Green, David I., 2012-10-22, A new fossil cricket of the genus Proanaxipha in Miocene amber from the Dominican Republic (Orthoptera, Gryllidae, Pentacentrinae), ZooKeys, 229, 111–118, 10.3897/zookeys.229.3678, 1313-2989, 3494006, 23166475, free, 2012ZooK..229..111H,
• Pérez-de la Fuente, R., Heads, S.W., Hinojosa-Díaz, I.A. & Engel, M.S. 2012. The first record of Protogryllinae from the Jurassic of India (Orthoptera: Grylloidea). Journal of the Kansas Entomological Society 85: 53–58.JOURNAL, Fuente, Ricardo Pérez-de La, Heads, Sam W., Hinojosa-Díaz, Ismael A., Engel, Michael S., January 2012, The first record of Protogryllinae from the Jurassic of India (Orthoptera: Protogryllidae),weblink Journal of the Kansas Entomological Society, 85, 1, 53–58, 10.2317/JKES111103.1, 83496373, 0022-8567,
• Hollier, J. & Heads, S.W. 2012. An annotated list of the Orthoptera (Insecta) species described by Henri de Saussure, with an account of the primary type material housed in the Muséum d’histoire naturelle de Genève, Part 1: Tridactyloidea. Revue suisse de Zoology 119: 149–160.WEB, An annotated list of the Orthoptera (Insecta) species described by Henri de Saussure, with an account of the primary type specimens housed in the Muséum d'histoire naturelle de Genève, Part 1: The Tridactyloidea (Caelifera) {{!, Request PDF|url=https://www.researchgate.net/publication/241277538|access-date=2022-01-31|website=ResearchGate|language=en}}
• Rodríguez, F. & Heads, S.W. 2012. New mole crickets of the genus Scapteriscus Scudder (Orthoptera: Gryllotalpidae: Scapteriscinae) from Colombia. Zootaxa 3282: 61–68.WEB, New mole crickets of the genus Scapteriscus Scudder from Colombia (Orthoptera: Gryllotalpidae; Scapteriscinae),weblink 2022-01-31, ResearchGate, en,
• Heads, S.W. & Taylor, S.J. 2012. A new species of Ripipteryx from Belize with a key to the species of the Scrofulosa Group (Orthoptera, Ripipterygidae). ZooKeys 169: 1–8.JOURNAL, Heads, Sam, Taylor, Steven, 2012-10-02, A new species of Ripipteryx from Belize with a key to the species of the Scrofulosa Group (Orthoptera, Ripipterygidae),weblink ZooKeys, en, 169, 1–8, 10.3897/zookeys.169.2531, 22371682, 3278811, 1313-2970, free, 2012ZooK..169....1H,
• Heads, S.W. & Leuzinger, L. 2011. On the placement of the Cretaceous orthopteran Brauckmannia groeningae from Brazil, with notes on the relationships of Schizodactylidae (Orthoptera: Ensifera). ZooKeys 77: 17–30.JOURNAL, Heads, Sam, Leuzinger, Léa, 2011-01-26, On the placement of the Cretaceous orthopteran Brauckmannia groeningae from Brazil, with notes on the relationships of Schizodactylidae (Orthoptera, Ensifera),weblink ZooKeys, en, 77, 17–30, 10.3897/zookeys.77.769, 21594149, 3088056, 1313-2970, free, 2011ZooK...77...17H,
• Heads, S.W. 2010. New Tridactyloidea in Miocene amber from the Dominican Republic (Orthoptera: Caelifera). Annales de la Société Entomologique de France 46: 204–210.JOURNAL, Günther, Kurt K., 2008-04-22, Die Tridactyloidea-Fauna Kolumbiens (Orthoptera, Caelifera) 'The Tridactyloidea of Colombia (Orthoptera, Caelifera)',weblink Deutsche Entomologische Zeitschrift, 41, 1, 1–56, 10.1002/mmnd.19940410102, 1435-1951, BOOK, Ricardo., Gómez L. de Guevara,weblink Estudio faunístico y ecológico de los caelifera (orthoptera, insecta) de la Sierra del Taibilla (Albacete), 1992, Universidad de Castilla-La Mancha, 84-88255-11-X, 435378980,
• Heads, S.W. 2010. A new species of Ripipteryx from the Ecuadorian Andes (Orthoptera: Tridactyloidea: Ripipterygidae). Zootaxa 2476: 23–29.JOURNAL, Heads, Sam W., A new species of Ripipteryx from the Ecuadorian Andes (Orthoptera: Tridactyloidea: Ripipterygidae), Zootaxa, 2010, 2476, 23–29,weblink 10.5281/zenodo.195303,
• Heads, S.W. 2010. The first fossil spider cricket (Orthoptera: Gryllidae: Phalangopsinae): 20 million years of troglobiomorphosis or exaptation in the dark? Zoological Journal of the Linnean Society 158: 56–65.WEB, The first fossil spider cricket (Orthoptera: Gryllidae: Phalangopsinae),weblink 2022-01-31,
• Heads, S.W. & Maehr, M.D. 2009. Proposed conservation of the name Gastrimargus Saussure, 1884 (Orthoptera: Acrididae: Oedipodinae) threatened by an unused senior homonym. Zootaxa 2268: 65–68.JOURNAL, Heads, Sam W., Maehr, Michael D., 2009-10-20, Proposed conservation of the name Gastrimargus Saussure, 1884 (Orthoptera: Acrididae: Oedipodinae) threatened by an unused senior homonym,weblink Zootaxa, en, 2268, 1, 65–68–65–68, 10.11646/zootaxa.2268.1.6, 1175-5334,
• Heads, S.W. 2009. A new pygmy mole cricket in Cretaceous amber from Burma (Orthoptera: Tridactylidae). Denisia 26: 75–82.WEB, PBDB,weblink 2022-01-31, paleobiodb.org,
• Heads, S.W. 2009. New pygmy grasshoppers in Miocene amber from the Dominican Republic (Orthoptera: Tetrigidae). Denisia 26: 69–74.JOURNAL, Song, Hojun, Amédégnato, Christiane, Cigliano, María Marta, Desutter Grandcolas, Laure, Heads, Sam W., Huang, Yuan, Otte, Daniel, Whiting, Michael F., 2015, 300 million years of diversification: elucidating the patterns of orthopteran evolution based on comprehensive taxon and gene sampling,weblink Cladistics, en, 31, 6, 621–651, 10.1111/cla.12116, 34753270, 53702892, 0748-3007, free, 11336/10677, free,
• Delclòs, X., Nel, A., Azar, D. Bechly, G., Dunlop, J.A., Engel, M.S. & Heads, S.W. 2008. The enigmatic Mesozoic insect taxon Chresmodidae (Polyneoptera): new palaeobiological and phylogenetic data, with the description of a new species from the Lower Cretaceous of Brazil. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 247: 353–381.WEB, 2014-06-15, The enigmatic Mesozoic insect taxon Chresmodidae (Polyneoptera): New …,weblinkweblink" title="archive.today/20140615150834weblink">weblink dead, 2014-06-15, 2022-01-31, archive.ph,
• Heads, S.W., Martill, D.M. & Loveridge, R.F. 2008. Palaeoentomological paradise: the Cretaceous Crato Formation of Brazil. Antenna 32: 91–98.JOURNAL, Heads, Sam, Martill, David, Loveridge, Bob, 2008, Palaeoentomological paradise: the Cretaceous Crato Formation of Brazil,weblink Antenna, Bulletin of the Royal Entomological Society, English, 32, 2, 91–98, 0140-1890,
• Heads, S.W. & Chesmore, D. 2008. New records of the slender ground-hopper Tetrix subulata (Orthoptera: Tetrigidae) from Yorkshire. British Journal of Entomology & Natural History 21: 243–246.WEB, New Records of the Slender Ground-hopper Tetrix subulata from Yorkshire (Orthoptera: Tetrigidae) - Research Database, The University of York,weblink 2022-01-31, pure.york.ac.uk,
• Heads, S.W. & Lakin, C. 2008. A new katydid of the genus Phoebolampta from St Maarten (Orthoptera: Tettigoniidae: Phaneropterinae). Polskie Pismo Entomologiczne 77: 256–266.JOURNAL, Heads, Sam W., Taylor, Steve, Swanson, Daniel R., Thomas, Michael J., 2015-01-30, Regional biodiversity of terrestrial Heteroptera and Orthoptera in southwestern Illinois, Technical Report Inhs 2015 (04),weblink English,
• Heads, S.W. 2008. A new species of Yuripopovia (Coleorrhyncha: Progonocimicidae) from the Early Cretaceous of the Isle of Wight. British Journal of Entomology & Natural History 21: 247–254 .
• Heads, S.W. 2008. The first fossil Proscopiidae (Insecta, Orthoptera, Eumastacoidea) with comments on the historical biogeography and evolution of the family. Palaeontology 51: 499–507.
• Heads, S.W. 2006 [for 2005]. A new caddisfly larval case (Insecta, Trichoptera) from the Lower Cretaceous Vectis Formation (Wealden Group) of the Isle of Wight, southern England. Proceedings of the Geologists’ Association 117: 307–310.
• Menon, F., Heads, S.W. & Martill, D.M. 2005. New Palaeontinidae (Insecta: Cicadomorpha) from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 26: 837–844.
• Menon, F. & Heads, S.W. 2005. New species of Palaeontinidae (Insecta: Cicadomorpha) from the Lower Cretaceous Crato Formation of Brazil. Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie) 357: 1–11.
• Heads, S.W., Martill, D.M. & Loveridge, R.F. 2005. An exceptionally preserved antlion (Insecta, Neuroptera) with colour pattern preservation from the Cretaceous of Brazil. Palaeontology 48: 1409–1417.

References

{{reflist}}{{BLP sources|date=October 2010}}

External links

• Heads Lab website

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