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Mediterranean forests, woodlands, and scrub

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Mediterranean forests, woodlands, and scrub
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(File:Biome_map_12.svg|thumb|right|upright=1.8|Extent of Mediterranean forests, woodlands, and scrub)Mediterranean forests, woodlands, and scrub is a biome defined by the World Wide Fund for Nature. The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in low-lying inland locations but can be cool near colder seas. Winters are typically mild to cool in low-lying locations but can be cold in inland and higher locations. All these ecoregions are highly distinctive, collectively harboring 10% of the Earth's plant species.BOOK, Cody, M.L., 1986, Diversity, rarity, and conservation in Mediterranean-climate regions, 122–152, M.E., Soulé, Conservation biology, Massachusetts, USA.,

Distribution

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- Mediterranean forests 1.JPG -
A Mediterranean forest, the Upper Galilee
The Mediterranean forests, woodlands, and scrub biome mostly occurs in, but not limited to, the Mediterranean climate zones, in the mid-latitudes:{{CC-notice|bysa3}} WEB,weblinkweblink" title="web.archive.org/web/20110401124536weblink">weblink 2011-04-01, Mediterranean Forests, Woodlands, and Scrub Ecoregions, World Wide Fund for Nature, 2010-05-27,

Vegetation

File:Ephesos amphitheatre.jpg|thumb|Close up of Ephesos ancient Greek amphitheatre with a maquis shrublandmaquis shrublandVegetation types range from forests to woodlands, savannas, shrublands, and grasslands; "mosaic habitat" landscapes are common, where differing vegetation types are interleaved with one another in complex patterns created by variations in soil, topography, exposure to wind and sun, and fire history. Much of the woody vegetation in Mediterranean-climate regions is sclerophyll, which means 'hard-leaved' in Greek. Sclerophyll vegetation generally has small, dark leaves covered with a waxy outer layer to retain moisture in the dry summer months.Phytogeographers consider the fynbos (South Africa) as a separate floral kingdom because 68% of the 8,600 vascular plant species crowded into its {{convert|90000|km2|mi2}} are endemic and highly distinctive at several taxonomic levels.JOURNAL, Cowling, R.M., I.A.W., MacDonald, M.T., Simmons, 1996, The Cape Peninsula, South Africa: Physiographical, biological and historical background to an extraordinary hot-spot of biodiversity, Biodiversity and Conservation, 5, 5, 527–550, 10.1007/bf00137608, This is equivalent to about 40% of the plant species of the United States and Canada combined, found within an area the size of the state of Maine. The fynbos and Southwest Australia shrublands have flora that are significantly more diverse than the other ecoregions, although any Mediterranean shrubland is still rich in species and endemics relative to other non-forest ecoregions.

Biome plant groups

{{more citations needed section|date=December 2017}}File:Matorral Cerro Aconcagua.JPG|thumb|Springtime in Chilean Matorral a few kilometers north of Santiago along the Pan-American HighwayPan-American HighwayMajor plant communities in this biome include:
  • Forest: Mediterranean forests are generally composed of broadleaf trees, such as the oak and mixed sclerophyll forests of California and the Mediterranean region, the Eucalyptus forests of Southwest Australia, and the Nothofagus forests of central Chile. Forests are often found in riparian areas, where they receive more summer water. Coniferous forests also occur, especially around the Mediterranean. Pine and deciduous oak forest are widespread across California.
  • Woodland: Oak woodlands are characteristic of the Mediterranean Basin and in California. Pine woodlands are also present in the Mediterranean Basin. California additionally has walnut woodlands.
  • Savanna and grassland: The California Central Valley grasslands are the largest Mediterranean grassland eco-region, although these grasslands have mostly been converted to agriculture. The remaining woodlands feature mainly oak, walnut and pine. The cork oak savanna in Portugal, known as montado, is a good example of a mediterranean savanna.
  • Shrubland: Shrublands are dense thickets of evergreen sclerophyll shrubs and small trees. They are most common near the seacoast, and are often adapted to wind and salt air from the ocean. They are called chaparral (California and southern Portugal), matorral in Chile and southern Spain, garrigue or maquis in France, macchia or gariga in Italy, phrygana in Greece, tomillares in Spain, fynbos and strandveld in South Africa, kwongan in Southwest Australia and batha in Israel. Northern coastal scrub and coastal sage scrub, also known as soft chaparral, occur near the California coast. In some places shrublands are of the mature vegetation type, and in other places are the result of degradation of former forest or woodland by logging or overgrazing, or disturbance by major fires.

Fire as a medium of change

{{Further|Fire ecology}}
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- Liebre Mountains 001.jpg -
Scrubland of the Sierra Pelona Mountains, Southern California
Fire, both natural and human-caused, has played a large role in shaping the ecology of Mediterranean ecoregions.JOURNAL, Gil-Tena, Assu, Aquilue, Nuria, Duane, Andrea, de Caceres, Miquel, Brotons, Lluis, Mediterranean fire regime effects on pine-oak forest landscape mosaics under global change in NE Spain, European Journal of Forest Research, 2016, 135, 2, 403–416, 10.1007/s10342-016-0943-1, The hot, dry summers make much of the region prone to fires, and lightning-caused fires occur with some frequency. Many of the plants are pyrophytes, or fire-loving, adapted or even depending on fire for reproduction, recycling of nutrients, and the removal of dead or senescent vegetation. In both the Australian and Californian Mediterranean-climate eco-regions, native peoples used fire extensively to clear brush and trees, making way for the grasses and herbaceous vegetation that supported game animals and useful plants.The plant communities in these areas adapted to the frequent human-caused fires, and pyrophyte species grew more common and more fire-loving, while plants that were poorly adapted to fire retreated. After European colonization of these regions, fires were suppressed, which has caused some unintended consequences in these ecoregions; fuel builds up, so that when fires do come they are much more devastating, and some species dependent on fire for their reproduction are now threatened. The European shrublands have also been shaped by anthropogenic fire,JOURNAL, Casals, Pere, Valor, Teresa, Besalú, Arnau, Molina-Terren, Domingo, Understory fuel load and structure eight to nine years after prescribed burning in Mediterranean pine forests, Forest Ecology and Management, 2016, 362, 156–168, 10.1016/j.foreco.2015.11.050,weblink historically associated with transhumance herding of sheep and goats.

Degradation

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Mediterranean cork oak trees (Quercus suber) in Monchique, in the Algarve region of Portugal
File:Royal_National_Park_NSW_2233,_Australia_-_panoramio_(23).jpg|thumb|A dry sclerophyll woodland in SydneySydney{{more citations needed section|date=December 2017}}Mediterranean ecoregions are some of the most endangered and vulnerable on the planet. As such, many have suffered tremendous degradation and habitat loss through logging, overgrazing, conversion to agriculture, urbanization, fire suppression, and introduction of exotic and invasive species. The ecoregions around the Mediterranean basin and in California have been particularly affected by degradation due to human activity, suffering extensive loss of forests and soil erosion, and many native plants and animals have become extinct or endangered.

See also

References

{{Reflist}}

External links

{{Biomes}}

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